optimisation de la gestion des régimes de débits pour les bassins versants de la Bourne et du Furon (affluents de l Isère)

Transcription

1 Parc Naturel Régional du Vercors Adresse postale : Chemin Fusillés LANS EN VERCORS Tél OUVRAGES HYDRAULIQUES : optimisation de la gestion des régimes de débits pour les bassins versants de la Bourne et du Furon (affluents de l Isère) OPERATIONS B4-2 ET B4-6 DU CONTRAT DE RIVIERE Tome I - RAPPORT D ETAT DES LIEUX DES MILIEUX ET DES USAGES Adresse Postale : 62, Boulevard Niels Bohr B.P Villeurbanne Cedex Tél Fax Portable : contact@asconit.com G2C Environnement Adresse Postale : Parc d activité du Point Rencontre VENELLES Tél Fax : g2c@g2c.fr La Bourne en amont du Pont Rouillard Décembre 2006

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24 $*! 9 I **D J!. 2!*! $H! *! CE $DB QE!H =, * 9&!* &;6> $DB = 9 c>. +"7 == $ 4 8 == 8? 4!H D J ' 9* E H!!E!H.4 = % *! 77L3++8!]9&+8!]$HH?I> %9 $ E$. 9 H!. ) B = >9 H 9! H 9$ =!! B >.) \ JH = * >9! H *!E =DI D! H 9DI * >. )!H!I! =/ %! >9! 9H 9 E! H B%.4 H % HI DI FE$J&++! $. 4!E HXD D!!H! $ $.' & ++!9. +"# F=A? 4 d"??9d"9d( +++=)"DR ' > $()', - HE!E* C!*!! J$ 4B =$'DR!! >9 $!D*! = (' >9 = H! > 9 $ 9!* $ =J $ D > H 9J$! O. 4! HE! E!!!! EE EE.4 $ D!!DB EE H J =*BDB 9 E 9 9!9[>9! D! H = * 9 * 9 [>! \! * D $ H. 4* * 4B B E BE! *.! "#$ %! % #

25 4 \ D!!E* 9H! 9!* H! =* >. % %$ C = %E >!B d"?? B & B H I DB*E 9 DE D E 9, $ B &7 8! $ 9D 9 / =! 4> \ D! d"?? B &.( &3D $ E ID E 9D E DE 9 B $ = H 4B > 4B (< B9 d"?? B &9 %!!! $I!! +++ C D* D % = H H! H9 D*> J D 9 H = H 9 % 9D*9* >9 *! 9! 4B = % > ( =J $ %!?>9!! D 9 HH!,, 9E HH! * DD 9! D! QE!E*.! "#$ %! % # 5

26 - 56&3 F!E* =A =A = - > = 56 ( DS 563 N B 56! +& '+3 " * " * " * " * " * " * " * d "! % +++&++ d"?? B & d"?? B & d"?? B & 56 ++&&++ d"?? B & 56 % -D! * * % DS ! -/ +;K+;K&;9 * ( +97DJH* B!E S ( 7DJ B!B! *!9 S F* ( 93DJ B!B D! *! % R 56 4 *? ( 796DJ B!B 56 DD H DD =, > ( &5 D J H B =E H!E* > 56 4 * ( 9&DJ B!B 56 O * DR ' D!9(' 9 % 4D 9(' D 9D 9 B9( B D 9D 9(< 9(- / % HI!!* D 9 9 9( = % E >!B N' D d"?? B & 4-4 ( 7 D J B!B =HI! E D > & D & D D = 9 < 9' B9? 9 B 9,N [> 5;& D 9 HI9 J I DE 9 *E 9!E HE =D!9 9 H[> 6 D N D! J I DE 9 DB*E 9 K!D* = 9! 9 9 H! > ;6 D N J IDE 9 DB*E 9!!!H =* H* J JH! > &+33 D HI9 * J I DE 9 *E 9!E 9E 36D!* D! JI DE 9*E 9 K!D*! "#$ %! % #

27 d"?? B & +53+ d"?? B & =A =A = - > = 9(< B9( - 9(< 19 % d"?? B & 94 9!*\ &&3+7 d"?? B & &&3&+ d"?? B & d"?? B & 56 (< B9(4 B 94 9 ( 9(\ -D (< B9(4 B9(\ B9( D! B9 ( B9 '* B9 B9(< %94 -?F?I 4 ( -?N2 I,!* 9 4B!* 4, N B B 4B ;;++++ d"?? B & 4D!* N d"?? B & d"?? B & 56 ( D 9(' 9( d"?? B & 56 (' N/ *B N'*?I D %* -H d"?? B & 56 4D ( B d"?? B & 56 9( 9(\ -D ;7 D HI9 * J I DE 9 *E 9!E!!!H =7 9 H H > 565 D J I DE 9 *E 9!!!H 9 K!D* =D!9\ > &5 D J I DE 9!!!H 9 K!D* =!H9 H9 > 655DJIDE 9E 9B. (! = *!* DR ' > 7&+ D D 9 9 9! D %JIDB*E B.&78! 9 D 9 *D 4 N I DE HE ;;D N JIDE 9 DB*E 9B.4 B B! E. D 9 H D* 9!*!! N'DR D 9 9!E9 HD J I DE 9*E 9 K!D* = R)R! > 3&D JI*E =JD 9 e 9 > 7D JI*E = C % 1! 6 " 6 31" > &;D HIJI*E!E =! 9 > 6+ D HI J I DE = > 5 D HI9 * J I *E 9 DE 9!!!E 9 DE =D 9 H > d"?? B & 56 9(' B ( 75D HI9*JI*E =D >! "#$ %! % # 7

28 - =A =A = - > = d"?? B & 56 '- d D! N' d"?? B & 56 '99( 563;++++ d"?? B & 56 D 9DD d"?? B & d"?? B & d"?? B )+;?6+&6& )&+?6+&6 )3?6+&; "?6+&35 "3?6+&3 +++ NI!! = > +++ NI!! = > +++ NI!! = > +++ NI!! = > +++ NI!! (\ -D (< B9(- 9 % 4 9(' 9 % 94D 9 D 9( B9( < B9(4 B9(- 9 D 9 B D 94-?F9( B9(< 9(4 9 (\ B9( D! B9'* B9D 9DD 9 B9 9 B 9(' B9 (< %9 4 4D 9( ' 9(- (9 \ 9 D D DD / 9\ D! *! I (-D I, *!* 4 9 9? N2 9 JD D*D % *! ( D*D %,!* 4 ( - JD 9H 4 9 9HI!E* D*D % &65 D D! J I D E DE ; D J I DE 9!!!H 9 *E 9 E = 9 B 9HH 9D!9D > &7+3 D 9 9!E9 HD J I *E 9!E 9 DE 9!!!E B =H!!H 9 D!9 9 D > &3D N JID E 9 DE D E =H H 9 D 9 > &55 D J I DE 9 *E 9B =B 9 > &5+; D HI9 * J I DE 9!!!H 9 K!D* =!H9 H9 9!9 4B!D* K > 557 D D =D 9 J *D > ;+ D 9 9 =* > &53 D 9 9 HI9 H H! H =D*9 DD > 575 D N!fE ND* N = H H! H9 %! 9 D*9 * 9* > &6;+D QE =B%9**9* >! "#$ %! % #

29 - "6?6+&37 d(+&?6&++&3 =A =A = - > = = > 9 O D % * +++ NI!! = > d '99( 4D 9( ' 9(- 9 O 9HI!E* D*D % ( / % HI! &+++D QE =HH > & D J I DE =5& NI!! > 4 8 F=A? &! "#$ %! % # 3

30 +"; / =!! % $ C E! =!! 4 FE$ HH U *, >9 E! = O 9)D 9 F D 9 4B!4 >9 = 9 - E! F J$!>9 =!!D!* 4> = ' 49 9 % B U,, U $9 B9 9 (? HH B>9 4 &;5+ B 9 9 $ 9H H 4B $ %!?. ' F =E &;;7>$I * ='> $!* 9 JH $".! "#$ %! % # 6

31 0 =? = 0"+ = 4 55!! R!!! *! %B. " C!!!! J$" 9 ) 49 )R! C!!!! 4 BB9!!!!. " %! 6!! B!! $ $! C ("-B9 ("' 9 (-' #?I 4 #?$2 9 (-' 4B 9 -H 9 9 J $" 9 E B B =! HD! >. 0"0 $ 5 8 = = F 3= ( ; &;;; $ 57++ D*!! C6L BB#! 9L B 1-9 6L BBR!.4 *! B 1 -! R!. )$ $ * -9!! E!! J $ = X *! $! $ >9!B H* \ D* =&+D*8!] 9&7J+D*8!] B>.! &;6 &;;;9! \!! =DS 9 9 (- (\ -D >!!! 9 F$D&+++ D* =4 9 P % $! 9 - U % &>. 9!!! &;6 &;;; =57L J (- >. E 9 *!! $D ='9 O 9-4 / 9 % / >.! "#$ %! % # ;

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

34 = O > - = - >. 7+++J&++++ J'9 J $! - 9 &+ +++ J J 4 =!! P! D*D>. 4! $!!H * $!!* E! = 9 9 E9[>. 4! J $ I * HH C gqbq9h DB E 9 DD JH $".2 H $ J( 9! J %! 4 9 <! B9 O =J $ O >9 - = ' - >9 H BD 9 H % % %!?9 B E!! DD?9 $. ( $*-= &;66J&;;>9 E %J * -! * I J.4 =&;;3J++>9 9 E! H! D!!* * - =!* &;;39!* &;;69 F9 V!* &;;;9 F!* +++9 F ++&9 V!* ++> 6&5H%! *. 4 =? $! J$ % 4B = ( = H >E!. ( H $*F H $!! * $ 9!! H 4ID 94$9 ()', ).44ID 9!* $ $ 9! H% *." I! D* $D! )!! EE, =),>.! "#$ %! % # 5

35 4"+ H= C09 < +9;5D = 540B6 $ & 44YID Z@67&;F&;6E!! J$ 9 C &K $ " 540B6 $! E H =&K+! *!B =! > $ &;69&K&+! &;69&K+! % &;6H % CD9DR 9! M6+! 5 K E *!! I * &K+! >. 4!!H $ 9 *!!*E I! &K&+!! $ FJ. $! J6+!hK9 $ H% *!!J! H E &K+!!. K $ " 540B7 $! E H! *. ( $ 9!! I E $ H *. %9 $E $H!* 7 J * =*! > $ $! 9 5K $ " 540B9 $! & $. 4"0 $= +6 = +990 ( 9 * *! $ *!!!! = H J $ 4 57 $! >.! "#$ %! % # 55

36 4"4 & %! 4$ C : 9 8$ 6!! 6 6 ; < :2! =! 1>? " 6@< 44$5F &;;9!H H &;;79! % EE H. HH! $ % 9 H. 4$ &+ 4$@;55F &;; $@;53 =! > ;535 = > ;! &;;5 *! F * *! HE EE =1-' 7 &;;= $ &7 4 $ %! $ $ * HH >! J H O$! H!! $ E *! $ J 6!hKD \ % ; &;;>. 4 *!!*E *HH $ J H FE$J! = &7 4$ $ ;3& >. 4"5 $= ; H &;;9 *! $ J $ $ DBE 9 $ 457 $!.' $ =* * *!! J $!!* >9 $ E I!!!! $ $. 4 $ %!$ $! $ 9!!!B *!!9!* $! 9 D%!B E!!."$! DJ %! $ *F! EE 9 C $%B %9! * 9 9J!E$J9E! $ $!9! E $ $! * 9! E $! E 9 $ D $ J$9 = I *>.! "#$ %! % # 5

37 4"6 %F C= 0 +7 = +9+9D = +9+9 = >!H $ 7 &7F &;6+. H!! C =7++Qi>9 H = $! EI *E >9 $9 Y Z E DBE = H% ) $>. 4"7 ' == 4$ 4555 $! E $ % $ ID! * 9 X!! $*! Y Z. 2 Y)!! EE, Z=),> *?)! ID 9 * ( ID." Y(D! )! Z = ()>! 7 *E E!! $! 9 *! J!. 4"# = & '? C'D 4$5F &;;9 ()',! H EE! % H.! $ E! $!! * H *F H E E $.) 9 *F H H%! E* *E 9!!! * $. 4 H * $H!.! )*-! E =)-> DBE. 4 )- *!!! 9 EE = ()',4 >.) 9 % H! $BI H ()',. ( * DR - 9 * *F H % H * E *F H E J!! JC H! DBE! * $ HH 9!!D %DBE! % % 9 $ = D*> H **E!!! E $! "#$ %! % # 57

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id. DBD 9 D $ * 9J J$D\+&+*F H $DB! ; 3; id9! # G6 idk. 4 ide! * HH E$ H!H.! "#$ %! % # 53

40 4"+, $= J #,B+6 C+5 K +9#,D # "!! %! H$*F H$!! *9 *E * 9 H!! $ H&;;7 ).4 %. * H $ *E 9 $ J %. H!! * 9E! $ $ % $. 4 5!&;;3 H =E *! >9! %*!B!. 4 * $ D J+! D =DD 4B > % E ) N" 9 D D N! =)">! R E!! *E 3+&7&V&;3+9 H 5!&;;3. 4"++ $= < ! =! $" 9 $" $! > ;!* &;;9 E E J $ E! HE 9! * &5 F &;;;. 9 E \ J %! E$J $ *!! DBE H % E $ % 9E * J$ % 9! $ HE \ J%!J $ 9E $H!J! $! H! $. 4"+0 H 00 > 0,,0 4$I H ++ $!! DBE H$! &7!* &;;. IE 5B \ * $I $* C \ %!!! 9 *! $ 9 \ $ HE 9 FE$ P $ % $ H 9 \ $9 9 P $!* J.! "#$ %! % # 56

41 4"+4 H >=? )! * 9 H $ H&;;7 )9!! $ $ I I H %!%.'9 I H %!! $ K? J E *! E$ \. 4$I H H +++ =E * $I H 5& F &;;3>! $ * < E $'* B.( I9 $ *! $'*= HH $> HC E gqbq $ HH J??O gqbq $*! 9 H!!* $ B!!!*??O gqbq9! H% 9 E B $HH J??O - $ J??O ( $*! 9 H!!* $ B!!!* $ % H! 9! H% 9 ID HHJ ID $*! $?)! ID - 'EE $" $)? ' H!!! $H! )?! % ID H J = OJ 3++!J$*! FE$* $'*>9 O. ( I9$! O D!* $ D. 4$IH ;* &;;3 $? * $ DB E ( 9H=FE$J&++!! >C E B $HH J??O - $ J??O ( $*! 9 H!!* $ B!!!* $ % H! 9! H% 9 ID HHJ ID $*! $?)! ID - 'EE $" $)? ' H!!! $H! )?! % ID H J = ID % >9 O.! "#$ %! % # 5;

42 4"+5 (= & 4 $*F HC % $ $ 9! % 9 R * J$! % 9! %EE \ D! 9 HH $ 9B!!E.! $H! C -"(E ))'?9))9( ))'((9 )"E 9 * (E9! 9!! 9 H *. 4))'? J $.4 ID! ID 9! EE.4 ID ))'?$!* *.! "#$ %! % # +

43 /'" C'"((')4' ((2'2 )(-"4"2: '12' "12( + = & +"+ & $ +, 8 4 H *!! H $ $. 4 %!! $DB! $= >."HH =* <9*! 9* N'*9* DD 9* $'* >9 HH H =* ( D!4B > = $, D 9 $ >. " %!* %!!! J E $DB! 9! J $! *.H9!* $ * 9* E HH 9 U $ $. 4 * 7 % E HH. +"0 = =? = ) *9!! HE!!! C!! DB E 9!! DB E, D 9!! DB E / D 9!! DB E D 9!! DB E,!! DB E B!! DB E JNN'* =(">!! DB E!! DB E (.! "#$ %! % # &

44 DBE!F * %. L / -B / L C>>D * < = %> ( B 2I$ M 0, 2 N 0, &3 ; + + O H & 3 + +? H ? H O & H 5& + + H * < => * DD = %> * DD => 5 & H " 3 & + E => BD H & ' H BD + + E / 4B 4B? * => = % > H * = %> => & & & H ; &+ + + D H 4B ; + + * N => ( = % > * N = %> ( => & ;&& & + H " & # %= =?! "#$ %! % #

45 >= 2= = == = B $ 4 2 %! FE $! 4$!! DB E! * <9 $ $! DB E 4C* < K5 $! C&K5 $! 9 H 9 H!! (< 9! )R! $!! $ )? )? 4 &;&7 % &K&+K&;;. 2 * FE$ + 4$*! $ C*!* E$!E =4j697+!9/j9;+!> C/j!U4j&7! %C $ 94 ++!9j;+++! 5 C $ ) U $! J! * =4j &6;!>D!* $ = 7+! 5 9 j;59&!,?> H 9! =4j76!9φj&9&!> L =B==C j78! C! )75& C E =&? & > $ >C! 94j66! $ &?C;79;+!,? $ C;97+!,? C! J ;97!,? )* * C&6K=&K&+!! > > FC *,!!! *.!!! $ %* C *! J! %&6K=FE$J++K> $EC * R!! =!I!!E $ >. E * H J = &++K> )* H! -D )* &C&96!5K *? C595!5K * C&7+K $C H!E!. * E! H $ ) =C+! (=? C37+Qi (=C! QiD 3C DE! O=C HE = $ > E C %H9 9 * H$ $ * *! * J! =! F >!, * $HH!!! 4 E! $ ++! 1 $ '!= J$!* > 1 $D* '! "!B?* = 9 >?* = > (!! H!!! 7 8 =! "#$ %! % # 5

46 >= 2= = B $ 4 2 %! FE $! )* )* H! -D )* 1 $ 1 $D* 4$!! DB E, D! $, D 9 $! DB E, D 4C $ $! C %! H 9 9 H (< C! H.!! $ )? )?! C* C/j!U4j797! %C! C& E $ U FE$J D!* $ * $! U $! 9J! * =4j5+>D!*! D H =4j+595!9φj&&;+!!> L =B==C j9&8! C! )75& C E *? $ >C $ &?CK $ C6+9+!,? C E * I M 7! 5 K9 H =!! +96+%+96+> * C! = $ 9 D J &;&;>. $C&57K > FCK $ECK &C&957! 5 K=H DBE, D > *? C597! 5 K=59&! 5 K ),> * C*!D J,j+9&3! 5 KU*!D J,&j+95! 5 K / C$ H H $ * 597! 5 KJ&! 5 K9 E H J&! 5 K= " 53>. ( J +93! 5 Kj+9;Qi9 ", D = &+>9 +95J+96! 5 K I E k5++k&+f ) =C&3! (=? C7355Qi= " C&37Qi> (=C&593! QiD!B 3CK O=CK E CK K!! '! '! "!B?* = >?* = > (!! H!!! # 8 = ' =! "#$ %! % #

47 >= 2= = == = B $ 4 2 %! FE $! )* )* H! K -D )* 4$!! DB E D D! *! 9 $ $! DB E C*! H DS C H 9 H. $ )? )? * FE$ +! $ * C* C/j&9!U4j56! %C4 &7+!9j++++! 5 C H L =B==C j5968! C! C E 7 $ 37+Qi 5= >! $! =,+9,9,5> $ >C % %9$!.4! % *,+,,&9,5, $ &?C&!,?U $ C!,?. CJ 396!,? D $ H=!* 9*!%!!j7+! 5 K> * C&73K > FC * $! =E GD! > ++! $ 9 D $EC R! DE!=HH > &C *? C9&! 5 K * C! / C!!!E! E * J &!=7!,?>U J! 9!! J+96+!U J! 9! 5! J+9+! ) =C5&! (=? C&;++Qi (=C) 3C) O=C E C JN.'H!*!% R9 H D. H!! J 9 D E * J&! 5 K!. H HD! =!! HH! HD* J$ $ >. $! $HH! K 1 $ '!= J$!* > 1 $D* '! "!B?* = 9 >?* = > (!! H!!! DS DD 9! (< ; 8 = =! "#$ %! % # 7

48 >= 2= = == = B $ 4 2 %! FE $! )* )* H! 4$!! DB E * D! * $'*9 $ $! DB E C* $'* H DS C H 9 H $ )? )? FE$ + C* 8/j9!U4j597! % 84 6+!9j) C H L =B==C j8! C! C E 7 &=?>! $ * $'*=,&> $ > C % %9$!.4! % *,+,,&9,5, $ &? Cc $ C+93+!,? C D!! $ HE! $ I *!.4!%!!! $ HH!! * C+973! 5 K > F C *! H D D $EC * 9 P D!!E! R =H HH!!! H!> & C *? C96+! 5 K * C / C N HD!* $ *$ ) =C&+! (=? C366Qi (=C) 3C) O= Cc E C D! = F D J! 9H$ Q! % > H HD*=!! HH! HD* J$ $ >. -D )= HH! * > )* 1 $ 1 $D* K '! '! "!B?* = 9 >?* = > (!! H!!! DS DD 9! (< 9 8 = =! "#$ %! % #

49 >= 2= = B $ 4 2 %! FE $! )* )* H! -D )* 1 $ 1 $D* 4$!! DB E! $ 9 $! DB E DS $ )? )? FE$5&K&K+6 C* C/j5!U4j7! %C4 7+!9j) C $ D %U $! D =4j37+!9φj93!9 j&.&+++> H =4j+!9φj&9!> L =B==C j&938! C! C E 7 &=8>! $ =,> $ > C % %9$!.4! % *,+,,&9,5, $ &? C&97!=&79+7!,?> $ C&97!,? C I = > * C= $ 9D J &;&;> > F CK $ECK & C * =J $ *!O D * > *? C6! 5 K * C&++K / C *H J&++K9$ H ) =C&&&97! (=? C637+Qi (=C) 3CK O= CK E CK K K '! '! "!B?* = >?* = > (!! H!!! DS DD 9! (< +, 8 =! "#$ %! % # 3

50 >= 2= = == = B $ 4 2 %! FE $! )* )* H! -D )* 1 $ 1 $D* "!B (! 4$!! DB E B! * DD 9 $! DB E B DD C* $ BC $ )? )? FE$+K+&K+7 C* V JI C/j79!U4j&5! %C4 67+!9j&9+7D! 5 =+93D! 5 > C $!! $F U $! D D =4j7977&!> H =4j35!9φj97+!> L =B==C j38! C! D H C E *? $ > C*D $ $ &?C;97!,? $ C5+7!,? C H9 D!E! J 5+79;U$*!%!! &7+! 5 K * C&! 5 KJ!! $ %= &K+!!!! $ %> > F C H $ECH!$D F DD & C&39! 5 K *? C! 5 K * C) / C&7&7!* 9 * F $ H H $.( I 37!.)&7!* &7CH! = *!! Q *E! e Q > ) =C&&+! (=? C7;++Qi (=C) 3C) O=C E C % D,R % e U* &;+! 5 K HD*! J 'HH! =H >J$*!! '!?* =*! ID >?* = D B>! H!!! DS DD 9 H* B9 * ( ++ 8 = (BB%! "#$ %! % # 6

51 >= 2= = == = B $ 4 2 %! FE $! )* )* H! -D )* 1 $ 1 $D* "!B (! 4 * $'*! $! 9 E $! % DB E C $'* $ '* C* 9 $! 4 $!! B ( B "9$ (B"!! =("> (B"!! =("> $ C % +5=IH )R! &7H &;;> $'* C % +5=IH $" F&;6> C!DBE % +5= 6!* &;55> C* * C/j&+!U4j3&9+! %Cj6+++++! 5 C $ $! J$!$ *! % eu!! J* 9 9*J D!* $ E $ * U % H =4j7!9φj&96+!>. L =B==C j&+! C!!! $'* B 75& C* i $ C $ &?C&;95+!,?= I F! > $ C * H E % = > 3C &> =1 j &7+! 5 K>9 > % $ (j&96! =!%.7+! 5 K>9 H! I * =! 9+!! > * C7++K=&K+!! > > F C JGDGH D $ECD & * JP * & C&69! 5 K *? C797! 5 K=3! 5 K G&6! 5 K > * C937! 5 K / CH! ) =C! (=? C&6Qi= &;6> (=C) 3C* O=C E CR H UD % $ JHH! HD*!! J$ = %D! >!! '!! =! > B '!?*?* = > =("c> +0 8 = &! "#$ %! % # ;

52 >= 2= = == = B $ 4 2 %! FE $! )* )* H! -D )* 1 $ 1 $D* "!B 4$!! DB E! * 9 $! DB E C* 9 $ $ )? )?! C* V C/j5!U4j597! %Cj&95D! 5 C $!! U =4 j++!>9d! $E* H $ &&++!. L =B==C j7++! C! Q! * C E $ *? $ C H 4 D 4B $ &?C7595!,? $ C735966!,? C! U *$!%! 7+! 5 K * C53K=&K+!! >J!! 7KJ D * 9! =J * H> > F C * D! $E 8 & C&95! 5 K *? C! 5 K * C / 8 H! ) =C7! (=? C6;++Qi (=C) 3C) O=C E C) HD*?! E!!,*! H E *! $* = E E H E$B > " J E H H 9 O * J$!! $* '!?* = > (!, 9.?*! HE ID. "!!E H* H!! =.7L H! > +4 =! "#$ %! % # 7+

53 >= 2= = == = B $ 4 2 %! FE $! )* )* H! -D )* 1 $ 1 $D* 4$!! DB E (?! * $9 $! DB E ( C* $ ( C $ )? )?! C* C/j&5!U4j7! %Cj7+++! 5 C $! 9 =4j5++!>9D! $E* H =4j&7+!9φj +9;+!>. L =B==C j78! C! J$!* ( 9 D? C E $ * $ C $ &? 8 ) $ C655!,? C*$ $% * &3! 5 K * C+K!! $ %=k&k&+!!!m&k+!! > > F C! $ $EC D $ & C&! 5 K *? C! 5 K * C) / CH! ) =C! (=? C;;+Qi (=C) 3C* O=C E Cc HD* '!!! '! "!B?* = 9 D ( > (!?*! H!!! $ +5 8 = 2! "#$ %! % # 7&

54 0 = > = 0"+ 0"+"+"+ 4 **DE $ * E =! **>C $"! D 9 $! D, D 9 E $/B QE -H 9 $DB 9D.2(( 9 DB, D U(. 9. 4! 9 )?#),9( 9 DB 4B J N =&Q!>=)?),> )?),.+.&;;59 DB % < =&6+ Q!> = 2'(( '4> )?),.+.&;;+9 * %I! J B =)2') '4>)?),&.+&.&;679 )?9 $! % H! * $'*. 0"+"+"0 4 $ E!!! *9 *. $ (= $ & $ i5++7+&+ )"DR ' 4? i55&7+&+ )"DR ' i557+&+ )? 44B i55+7&+ )"DR ' 4? 4 i55++&+ )"DR ' 4 4 i55+&+ )? 4 (< % i5577+&+ )? 4 B +6 8 C )D! "#$ %! % # 7

55 !! 9!! $ 3BB / (BB % B B1BB $? # +97; #, +9#+ +9#0 +9#4 +9#5 +9#6 +9#7 +9## +9#; +9#9 +9;, +9;+ +9;0 +9;4 +9;5 +9;6 +9;7 +9;# +9;; +9;9 +99, # +99; ,,, 0,,+ 0,,0 0,,4 ) ) ) +7 8! "#$ %! % # 75

56 0"+"+"4 &? 4 % HH 9J)? ("9!!E!* $ %!! $H H! $. 4! )? C 4B? I ( P 4 < 4, D 4! '*, DD - &;; &;; &;; &;;7 ++ < ++& ++ - &;;7 ++ < ++& ++ - &;;7 ++ < ++& ++ - &;;7 ++ < ++& ++ - &;; &;; &;; &;; &;;7 ++ ) & < ++& ++ < ++& ++ < ++& ++ +# 8 &? 2 C, D C E F ++&++9 C E F ++&#++. 4! (" C )*F $ ++& ++9 E! J$ J'*. $ ++ P = P +99; C Q 0 D 4 * HH 9E?! DBDE H H! QE." H $! $ = %! >9E D! J HH = 4 %! >.! "#$ %! % # 7

57 ) % DBE 9! ^ $! * J! $!? H J HHN!DE! D! C H C D H9 Q! %9 H H C $! QE *.! HH $HH DB!E J! F! 9!! J!!E *! * $ % H O. " J E 9 $E $! DBE *?9 $H E$ *! J!D D! J$!. %*Y Z %*Y ZE!!. 0"+"0"+ $? & >= ) $E $ 9! $ $ I!.)HH! EH C )=*E $ >C*F &+F9 1=*!!>C*F FH9 1-'=*!!! DE >C!B *F!$ = >9 1-'7C 1-' EN YEN EZ." $ *!!! B **&K7 I. 4 1-'7 H =H! ;535;!&;;59 4N5F &;;>.(! $ B E =HE >. $H* E. )$9D * 9 HO D9 C! E!! HH O * H E$H. H9 E!! 9! ) 1!*. $ %$)?H! D 9 E!* HH! HE $. X D *%< &;3J++= $ Y/B % <Z>.' B E! %!!.! "#$ %! % # 77

58 P / ' 1-'?E!?E =! 5 ' K> %! T %! T =! 5 K> %! T %! T &;; &9+ &;;3 & &9+ &;; ; &;;6 &9; ; ++& & 59& ++& & & & +; B0,,0 B F= T4HE %!! H! = + P $ $D. 4 D =! > J $ E # H J+.7. " $B DE =&;;7++9* &6>E *! $ H* 9 &;;69 &= && &>.4$ &;;6* D 7 =*.&6>. ' 1-' =! 5 K>?E %! T %! T ' )*!B =! 5 K> /?E %! T %! T &;; +9&6 +9+& 3+9 &; +96; +9+& 3+9 &;3 +9&; +9+5 ;9& &;; +9; ;9& &;7+ +9&; &695 &;3 &9&& &695 &;& +9& &59 &;75 &9& &59 &; +9& +9+;7 &+97 &;73 &9 +9+;7 &+97 &; +9 +9&&7 693 &;3& &97 +9&&7 693 &; &5 39 &; &9; +9&5 39 &; &7 9 &;7 &95& +9&7 9 &; &3 793 &;; &95 +9&3 793 &;7; +9 +9&; 79& &;7; &97 +9&; 79& &; &3 9 &;3 &93 +9&3 9 &; &;7+ & &; ; &;6 & ; &;3& &;3+ & &; ;6 59 &;& &9& +9;6 59 &; &6 59& &;77 &95 +95&6 59& &;; +95; &;;3 & &;36 +95; &;3 & &; ; 9 &;7 &9; +953; 9 &; ;; 97 &;5 &9; +95;; 97 &; &; 9 &;3 &93& +9&; 9 &;; ; 95 &;7 & ; 95 &; &;67 & &; & &;6 & & &; &;;6 &9; #B+9;# R +996B0,,0 B F=! "#$ %! % # 7

59 Débit mensuel d'étiage de la Bourne aux Jarrands (m3/s) 1,4 1,2 1 débit en m 3 /s 0,8 0,6 0,4 0, ( &;;7++5 ( &;3&;63 Débit moyen de la Bourne aux Jarrands (m3/s) débit en m 3 /s 4,5 4 3,5 3 2,5 2 1,5 1 0, &? 1! "#$ %! % # 73

60 ( E ; =* &;>9 &;;3 &;;6!!! D.( 9 $ &;;6 X!!! =E *!B K %>.(*!!! *$ *. $= = == & > >= "! E 6 HO D! 9!! * $ E. $& = = & +99;A F= & 2" D% E $! H J $ H C $ &;;6 HH Y H Z=! 9*$ * >9 *! $! F! $HH = >.! "#$ %! % # 76

61 0"+"0"0 = L 4 * %HH H H!* =! 9 H9 HH!! 9 [>!! J!* %! %!E. 4! $!*!! * 9 E$!* = HH!E 9* E $DB >9 $IC *!B O HH B H. ' HC %HH C- 9)D 9 94B 9 %$ QE C, D 9, 9 $'*9 9 %!! DBE C $ * ! "#$ %! % # 7;

62 -J L & S -S $ <!, D!,, D H )D! <!, D!,, D H )D! ( N'* ; ( N'* N'* &+ && & &5 & &7 & N'*, DD B H $'*, DD B H $'* H 4B &3 H 4B H " 4 *J$! < Y Z! DE $ U * O9 <9 %? 9 O -.4 * DE %<!J&3+8!]=* &+8!]> 4, D! QE U H E$ )?=!!! 4J H$! * >! E! =j*> * E! =H >.4 O $ 8! )! J! O * DE $ &+ 8!]! J \ $!,. 4 *, $ 3+8!] 4 $!, D $ 8! 4 % 7++! 4 * J! 58!] 4 $'*! $ QE! % U!!H 4 * * $'* 8!] 4! QE!I! B! E $'*.)HE$ )?! E! =j *> * E! =H > O! E E E \! =! H H > ' * $'* $ U!! J$ $ J '$ O! $'* $! =!!! J$ > 0, 8 F! "#$ %! % # +

63 X * OS %B!! $)? )"=! *E /B>C *^+ &;3J++=! F J$!* <>9 *O&& &;;7J++=*J$! DD J $ %$)?!! >9 *O& &;3+J++=S J! - >.,S J 9 %E! C = => = =5> B! $E9 X! HH OC!* $ % U Y Z J **D. 4. 0"+"0"4 F & 4! O! E %9J!! $)? (". 4!! )? $DB C $ J *. C %* <9! $'*9! H YH $ Z $H Q $. D * 9 C = ** * D $.4 * E! %<$X *!!. % $, D 9! * C H$ % J$D *! E $ &;&;.X **! "#$ %! % # &

64 S % $ % $)?. 4 *! = > * D. DD E J H Q 9 $ B H.'! I $E C = *!9 * **! )?. 4$!! ("! * $'* 9 E!! DB E.4! E $ J =E % E$ J >.4! H* =! >C H $ 9 *!.!!!! *. 4 * * $'* HH JX. HH 9 * * 9 $!!! $" = $ - > H! $ B. 4$ E % $ $'*!* $ % $*E! $ * * = * I!! E H H! >. 2*! ++&C! E % 9E!! H* * &;;6= % E H! >.4 * $E$ U * &. 1 DE =! 5 K> 1" 2" /" " / 1 1" K " =" -" =" / & && & ;& = P $ P +99; 4! * ("! HE$* 7++KI! J$* $'* U$ E H C =! "#$ %! % #

65 0"+"0"5 I.&...& 4( " 4 *, D * * =, D >9 * = H* D!* $ E $! = L * *>!. 4$! *, D HH! QE!U X *$! J."!E # $ D % $ = % -D 9 HI 4* > U H! J 7+8!. 2 *, D ), $)? =( > &;;; $ Y/B, D ZJC $! **DE 9 $ % = &;767;9&;336& &;;7;6>9 * J $ =!>. 9 $ $!!! $9 * * *.! HH! *J, D %<.* C 1 / B 1 1 B = B = 1, D =! 5 K> &9& &9; +93+ &957 1! <=! 5 K> ;6 &93 HH +L 65L 3L 3;L 00 8 ' = C E HH!!B *! * $ &;;69 E *, D < $!I!! 9 E! $E HH! % *!. * *, D &;;6 DE C 1" 2" / / 1 1" K " =" -" =" 1 DE =! 5 K> &9 &9&7 & &9+ & & &9;?! +9T1 <9 +9T1 <9 +9T1 <9 +965T 1 <9 +965T 1 <9 +965T 1 <9 +965T 1 <9 +93T 1 <9 +93T 1 <9 +93;T 1 <9 +93;T 1 <9 +93;T 1 < ' = CD P +99; CD! "#$ %! % # 5

66 ' =! DE H! H! $!! C B! QE HH $ J $ HH B!E $! 9!!, D 9 $! H 9!!! %<. ' NF HE, D!!! H E! %9HE $!B * $!.4$! **! E! )? *! F. %H!! *. H! $HH DE *.4$E E C = $* * > 'H 9!! * J $!, D C * H C = 1" 2" /" / 1 1" K " =" -" =" / 1 =! 5 K> &9 &9&7 & & & &9; &9; 05 8 ' = P +99; CD 4!B! H! % **D..&... 4, ' $ %! DBE, 9D!E E H!**DE CE * $ 3+8!], D."!H *H $!! =&7+!!KJ'>. (* H!* 9! * *,'0! $ $! D, D. '!B 9!I!!D E!! C I * *!!B, D, U HH E%*!, D &;;6H$ %,.! "#$ %! % #

67 4$!* * %7 C 1 / 1 1 B = B = 1, D =! 5 K> &9& &9; +93+ &957 1, =! 5 K> 9; 976 &9+ &9;5 HH &33L &55L &;L &5L 06 8 ' - 1 DE =! 5 K>?! 1" 2" / / 1 1" K " =" -" =" / 9; ; ;5 9; &96 9 &933T 1, &933T 1, &933T 1, &955T 1, &955T 1, &955T 1, &9;T 1, &9;T 1, &9;T1, &95T 1, &95T 1, &95T1, K 07 8 ' - P +99;.&...5 O, D,, D, 9! 9 HHU * &+Q! 9 $ J\ $!,. 2!! *,'0! $ $! D, D 9 * *HE, D.! "#$ %! % # 7

68 !!!! = *! 9 HH %*! >9! O$ &;;6C 1 / B 1 1 B = B = 1, D =! 5 K> &9& &9; +93+ &957 'O5=! 5 K> ; +9& +9 HH +L &7L &3L &L 0# 8 F L ' = B ' - 1 lo5 =! 5 K>?! 1" 2" /" " / 1 1" K " =" -" =" / & & T 1, +9T1, +9T1, +9&7T 1, +9&7T 1, +9&7T 1, +9&3T 1, +9&3T 1, +9&3T 1, +9&T 1, +9&T 1, +9&T 1, K 0; 8 %= L ' = B ' - P +99;.&... 4)D 4)D HH E $H!.)! % = J 7.&.&;6>9),N)?! * )D J &8!]. $* $ 9! * )D E$ * %<9 * H=&8! )D &3&8! %<>9C * H C = / = 1" 2" /" / 1 1" K " =" -" =" 1 =! 5 K> +9 +9&6 +9& & +9& &; &7 +9& %= = P +99;.& ( '* 4 $'* = > =! > J!I! QE!! 4 D $'*9 H Y Z. 9 *9!!!H!?.! "#$ %! % #

69 4 $'*9 9!!* J*!B!hK * +!hk. 4 D$'*9! $'* 9! E! D! *!=+!hk>. 4 = C&6!>9! 9!! E E E H 9! *! 6+!hK.! '. S J! * D. 4 I!! \ B $'* $!. 9 H! J!! D $ %! * $'* J&;8! $'* 9!! $ E*!B +9!hK.4 H 9 D $ *$ 7+!hK= 4 >! D $* ++!. 4 E! 9 * *9!! $! \ B.( $! =<.<., 4! 9?D '&;36>9 \ B %*D C! % $'* % = 4 >. 4 4 H!!! D! $E* 9 %! 9 9! % $'* =D$'*>. ( % % % *! $ *9 $!!.)D 9 H! ( (< P % H!*! H. '$D 9!* E $'* E 9 E!H HO C * $! H $'*9! H* 9 % $'* H U * % J H * H9 H 9 $'* ** * J E U!! '*9H %. = & E $'*!!!H 9 $ %! *9 $ E E F $* E $.'9/. H! * $'* * J$! < $ &;5. H H! E J $ &;;6 H* $'*=!B 97! 5 K>9!* $! =$E$X! DBD B!$* >.! "#$ %! % # 3

70 ) * * *DBE!! * 9 * * $'*!! $E=>* D. * H C = 4 *! $'* $ &;;6 * C = / 1" 2" /" / 1 1" K " =" -" =" & 1=! 5 K> &9+ 796; 9;; 9&5 &9;6 9; ;5 4, 8 3? = & P +99; 4 * *!! HE *!! C %! YZ ( $'* 9 E % $!I! U $H. = 4! 9H9!! $D9! % % QE 9 E %!I! %I!!! %. "!*! $! = * >9 = * $DB9!!, D %! >. 4 * $ % $)? E % $!, $ $! $ *. 4 D $ * FE$J D 6+!C! %!! * H! 1j!4D D=!j+9567 H!H )?>. (! ++& ++I!J C E * 9! * J H E$! I =H >. C! 9 HH! %* 9 J! % $! 9! QE! % %.! "#$ %! % # 6

71 2 0 8 = 0,,+ 0,,0 2!D!E *! % <9 * % 9 H. HH 9$ * J $ &;;6 * H % * *. H 9 $* D 9 $DBD YJ!!Z C.&... 4 = 4 * H S J F % )? B. X * &;;6=* *E DB>C / 3 1" 2" / / 1 1" K " =" -" =" 1=! 5 K> &97 &9&7 &95 93 &9& +9; ; +9;5 &9& &9& &9& & P +99;.& B 4 * 4B $ %C! "#$ %! % # ;

72 ! $F! * H UH D B! QE 9X! * $! $! * J!E H* 9! %4B C%( 9 9 J (< B. ' $ J $D. 4 H C * => CTUD CIIT 0 D * CVID 4 ( &;&&; ) ) &;;&;6+ & 793 &9&5=) > (.= HH!B! >. 9! * HE H! E *! %* H %QE. H $* $ 9 $! * 4B J H % % F C D (. E *JH J % J B * HH j&k57.' *DE H ++ K ( D! B (" H J = $'* 9F! $ >. H 9* C 1" 2" / / 1 1" K " 1 '* =! 5 K> =" -" =" / 593 9& P +99; CD H D ** J! - =P *$! H J % 4B >.' H* * 4B!! $E=5>U* C #$ =!" 4! +9 %++ * ( D! B (".4 C 1" 2" / / 1 1" K " =" -" =" 1 '* =! 5 K> / &9+ 79& 9 & & P +99; CD! "#$ %! % # 3+

73 .& H H F % QE." %I!! HH \ $H9! %. ( O! H! J F C $ 9!$ * $I!. HH X! %H =D &.> UE$ - 9 * =DE > * f C * J 9 E! Y H! Z B!. )! 9 H * H!! % * =!> = *! >C I *! HH J.' ( $'* I %! 9$!!E *!B O %D %. ) $E 5 * D9 * C!!! C =!" #$! "#$ %! % # 3&

74 0"+"0"6 % 8 = 4 * EB! * DE O.!. -J L + ( & S -S <!, D!,, D H )D! <!, D!,, D H )D! ( N'* 1 + j1!< 1 & j1 + 1 *.< j1.< G1.< 1 j1 & G1., D 1 5 j1 G1, G1 O5 1 j1 5 G1 *., D 1 7 j1 G1 *.< 1 j1 7 G1 )D 1 3 j1 1 *.4! 6 ( N'* N'* 1 6 j1 3 G1 $'* ; N'* &+ && & &5 & &7, DD B H $'*, DD B H $'* H 4B 1 ; j1 6 #1 *.'* 1 &+ j1 ; G1. 1 && j1!dd j=1 &+ G1 *.4! G1 *.'* G 1 *. > 1 & j1.dd G1.DD j=1 && 1 *.DD G > 1 &5 j1 & G1 *.DD 1 & j1 &5 G1 G+. 1 &7 j1 & 1 (" & H 4B H " 1 & j1 &7 G1 4B j =! "#$ %! % # 3

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89 & =>= 3? > >> (BB%! "#$ %! % # 63

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99 $ % =' % ;>. D D $* H*!E = HH H> =*! >. 4"5"0"+ 4 * * B ( ID $' 3. ID E = 9 9ID > &;65 ++&. H9 $ * =!9!* H! > I! HH H *! = H >. $ B E! H!9JH$ = >!= >. ) F * 9 J 6 9J+ 9&+ * 4B 93 * 9 -?. 4 E *. ($!* * =6>9.!! 9 =&;;+> 9H H $! = J$ * DD ' >. 3 9,. B!9<..=++>(BD &59 %HH. ( ID '&69.;G %.&;97! "#$ %! % # ;3

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

102 $7 + (! = 4>9Y!Z ' E &;;5 ++5.! * $! J $H DB E. E DBE HH = $ \ >. I!! H. 4 H D* % 9 E! ** E H*B %. % 9 4 HH H *%!! *! = 6+++KD9H 7>!I! %I! \HH =!!! 7+++KD &;;!%!! &7+++KD +++>. ) 9 F $ 93L=&;;;>J9L=&;;3>!B $ +L $ =H >.4 + G $D 7+=+ G >KDJ ;+++=+ G >KD %!I!. $ I *9\ F %. Effectifs de 0+/Effectif total (%) Truite fario (0+) - Station Pont des Asniers Densité Proportion Effectifs (N/ha) 22/09/ /09/ /07/ /06/ /06/ /06/ /07/ /07/ /07/ /06/ < & C, R D > ( = 8 $ (H= " E J $! + G H*. HH 9 B D =!*!I! >! E!!! %!. H $!* * 9 % H D &;;5=* >. %=& G > = G > DHH D E 7L 5+L.! "#$ %! % # &++

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

104 !! 9 H* HH H + G * &;;;! E H* 9*!=7 3+! 5 K H!>E$J$! = &5+! 5 K * >."$ * H! H* 9H E $ ++E = F %!* >.! F=769! 5 K! 3F%<9579! 5 K DD > %E H* = > + G *! F!I!. 4$ HH! 9!! $B H + G! E$ DBE E $* ^H! E $! =9&;;79)H >. 4 *!! *!!I! E HH H!!!* &7QKD=++>!%!! 5+QKD &;;5= ++ QKD +++>!B $ 7+QKD. H = HH H *! >!* % H $. " E 9 SD \HE 9 9 %E.$!! $ +++E *!! 9EI D SD E H E E! J $ $ =- >." E$ $ * H! E.) 9 9 $H %. H9$! ID B!! $ J.' E &;; +J5!$!*! $".! "#$ %! % # &+

105 Biomasse (kg/ha) Truite fario - Station Pont des Asniers Effectifs Biomasse Effectifs estimés (N/ha) 22/09/ /09/ /07/ /06/ /06/ /06/ /07/ /07/ /07/ /06/2003 Biomasse (kg/ha) Truite fario - Station Camping de la Balme Effectifs Biomasse Effectifs estimés (N/ha) /06/ /06/ /06/ Biomasse (kg/ha) Truite fario - Station Colombier Effectifs Biomasse Effectifs estimés (N/ha) /11/ /10/ Truite fario - Station Passerelle Béaudet Biomasse (kg/ha) Effectifs Biomasse Effectifs estimés (N/ha) /11/ /10/ /06/2003 Biomasse (kg/ha) Truite fario - Station aval Lyonne Effectifs Biomasse Effectifs estimés (N/ha) /07/ /07/ /06/2003 Biomasse (kg/ha) Truite fario - Station Pont de Manne Effectifs Biomasse? 6 8 > C>>=> D 7 B = 8 $ (H= 17/07/ /07/ /06/ Effectifs estimés (N/ha)! "#$ %! % # &+5

106 4$B * '! E HH H %!* 9 9!! $B 9 * H* J5!.( =&;;5++5>9 HH HH HO!. * \ M+! &;; &;;9E$! +++9 J D! HH E E!. 4 HH H %!! ++& $!* 9!* J* Densité d'individus (N/ha) Truite fario - Station Pont des Asniers >20 cm >23 cm juin-93 déc-93 juin-94 déc-94 juin-95 déc-95 juin-96 déc-96 juin-97 déc-97 juin-98 déc-98 juin-99 déc-99 juin-00 déc-00 juin-01 déc-01 juin-02 déc-02 juin = C0, = <& +995A 04 = = D ( = 8 $ (H= 4$ HH ID HH J! DE D!! &;;= +J5!> E$ * 9 HH $.!9! I D $!* $! + 5!. 4 JH. Proportion (%)/ population totale Nb individus/ha Proportion Nombre d'individus /ha 22/09/93 28/09/94 17/06/95 03/07/96 19/06/97 17/06/98 11/06/99 18/07/00 11/07/ /07/ /06/ # 8 CID 0, 04 = C "! "#$ %! % # &+

107 "! E! J $ &;; P D &L9! 7L!E."!I! B $9 E H!!!!! &;; KD9H +++= SD c>.9 *!! E E HH $! E '. 2" D*9 HH HH HO! 9 H * *! HH H &;; &;;=E $ 9$ $ 9H 6>.4$* $ &;;7! D *.4! 9! HO! 9 &;; &;;;!* *! E 9F! * &;;5 &;;. ( *!! ++5! J * &;;5. 4 D* * E % E $ $D*! *!. ( H! BH!! = 9 E H $ J $!9[>.)!I! HO9 J$E* =!J!* J * *>! H!. 2 ) 9 ' 49 H!9E 9J 9 H! $ H!! DB E E$ $B F HH. 4 * ' H 9 FE$J 5+QKD =&;;5>9!!* I H!!H! E! H! *!.!9B!E!* HO!!! H $DB = > $! HH H + G. S9 *!! = H 7>!I! I =!! c>h. HH 9 E! 9! 9 E! H B!E.!H!! H $ R E ID E.( % =! > B *F H $!!!DE =!! >! $. 4 *!!! HH ID 9 $! $ * *.! "#$ %! % # &+7

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

109 ( $!* 9 X E* 9! H E* F $ =+ G > HH H! D &;;3.4 H* + G * * 9 E! &;;;. $-= +1 9 ) 9 = >9 H &;6+ $.' E HH $9 H 9$* D*! BH! B! =*! E $ % F $ 4>. ' 9!-J! = % * $! 9 J / *! >9 H D* = &;; &;;6>.44D HD $ E$ &;; &;;3 E!.! ** = E H *B >9! *. 4 HH H H +++ KD *! H J7+QKD=H 7>.) HO \ 9 + G H* &;; &;;6= &L> &;;3= 6+Lm>. DHH C * HH H +G9 H* &;; &;;6 = 5+++GKD> H &;;3= 5++++GKD>9 H* S. 4 H &;;3 $ %E!* %! = * $! *! 7!>." E + G!* J * ' 9 HH *! = &J&+!! &;; &;;39 $J &J! ' >. 4$B D!! HE! E!! E Y!Z9 $! $ HXD. E $!= > % 9 H*! %! =+ G J& G >! H* * = H J 7!! F>! D!! E! D K*!. 4! H 5+L=D &;;3> &37L=D &;;>9DHH E H$! $S &;; &;;3. % =& G J G >9 %! 5+ 7+L. " * E J $ 9!!!!E H*! + G = HH H!> *! F $ H ID. ( 9 D* * = > - J $ < =, D >. 4! "#$ %! % # &+3

110 ! J $! E $ 9 H!* QE =, D 9 >.4 HH H! %I!! H* =! 5++KD>!I! E *! =H J3QKD>."!* B *! &;; =$P HH H!E >9D EY!! Z FE$ &;;3=*! >!E!* * H. 2 4 /!H! E DBD!E $ 9!! * D! * =E *>."!* E!! %HH * < =, D 9!! >9 * *! E $ 9 E!* H! B DB*E! * =",9 >. H9 * $ * < = F ( 4> E! J$ E 9*. 4 H! E D* H! H.)$ (9 $D** H* = H > E * E!.) 9!9!E = J!* D HD* >!! $ D!. H9 $ * =9 9 H *>9!I! 9!! *! E E E. HH 9 = J $E* > $ B!! H*.! "#$ %! % # &+6

111 Biomasse (kg/ha) Chabot - Station Pont des Asniers Effectifs Biomasse Effectifs estimés (N/ha) 22/09/ /09/ /07/ /06/ /06/ /06/ /07/ /07/ /07/ /06/2003 Chabot - Station Camping de la Balme Biomasse (kg/ha) Effectifs Biomasse Effectifs estimés (N/ha) /06/ /06/ /06/ Biomasse (kg/ha) Chabot - Station Colombier Effectifs Biomasse Effectifs estimés (N/ha) /11/ /10/ Biomasse (kg/ha) Chabot - Station Passerelle Béaudet Effectifs Biomasse Effectifs estimés (N/ha) /11/ /10/ /06/2003 Biomasse (kg/ha) Chabot - Station aval Lyonne Effectifs Biomasse Effectifs capturés (N/ha) /07/ /06/2003 Biomasse (kg/ha) Chabot - Station aval Pont de Manne Effectifs Biomasse /07/ ; 8 $ C>>=> D 7 = 8 $ (H= /07/ /06/ Effectifs capturés (N/ha)! "#$ %! % # &+;

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

118 (!!DE =!* %! > H! H* E J $! HH * E $ %!."!! H D!E! * $'* (\ B9H E$ 9!* D! =H > E $!* ( $ 9E D &;6 &;;6. (! =! >9!! &;;3 9J 9 H 4D HD.' $* D*9 EX!! E $ E!* F! B. 4 D*! * &;6;&;;+ % $ H - = <, D >. $ E 9,.4!* I J*! E E $. 4 9!! ++ QKD H HH H!! F $ =;L + G >.! H! E!9 $ $* D* $. = J $!,9 N J J $!! - >9! DJE &;; &;;6."! E 9 J 9$D J &;;=+9QKD>.H9 9* $ &;; *!! &;;=396QKD> &;;3=&597QKD> * &;;6= &57QKD>. Nombre d'individus/ha CHE TRF Nombre d'individus/ha LOF VAI C>>=>ID & ' C/D C%2D $ C$)D CDA 3 C3D = >= C2D CD" '$ 9! H!.4 &+QKD &;;J! &QKD &;;69 *! 4D 9 +QKD=&;;>J &+QKD. $! E$ &;;9!B = &++8KD> * J! KD &;; &;;39 * H! J 659QKD &;;6. 9 &;;&;;39!!* J * '!. &;;3 &;;6E 9 *! $! E HH H!! "#$ %! % # &&

119 !*.! H $ $ B! $ H* % &;;3 &;;6. 4$B D!!! &;;6 H* * $!* $S F $ =+ G >E E 795L =5;9KD>.)$ HO 9 + G H* E$ 79&L=&;;>J795L=&;;6>9 E! *!." $ * H!!! HC E! * $ =FKF >9 J E =k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

121 Biomasse (kg/ha) Truite fario - Station amont Pont d'echevis Effectifs Biomasse Effectifs estimés (N/ha) Biomasse (kg/ha) Truite fario - Station aval Pont d'echevis Effectifs Biomasse Effectifs estimés (N/ha) 28/09/ /10/ /09/ /09/ /09/ /09/ /10/ /09/ /09/ /07/ /09/ /09/ , 8 > C >>=>D & & CD &=" Effectifs de 0+/Effectif total (%) 80% 70% 60% 50% 40% 30% 20% 10% Truite fario 0+ - Station amont pont des Echevis Densité Proportion Effectifs /ha Effectifs de 0+/Effectif total (%) 80% 70% 60% 50% 40% 30% 20% 10% Truite fario 0+ - Station aval pont des Echevis Densité Proportion Effectifs /ha 0% 28/09/ /10/ /09/ /09/ /07/ /09/ /09/ % 28/09/ /10/ /09/ /09/ /07/ /09/ /09/ < & C, R D > 3 &= 4 HH H + G!! * =H &&>.4 $D! 7+KD=&;;79&;;69++&>J 6++KD =&;;3> &7 3+L=&;; &;;3>.4 Y! Z9! H*! * E 9 * E ' =!>. 4 H* &;;6! = $ E E *! E >. $! 3"! '$$D 9 E!!* J!C H. 4! * = D*> H* = 4D HD >.' E$ ++&9 HH H H* E % E 4D HD!* HH=55+KD>. H9 HH H B!E!!E!C $D +++KDJ &7+++ KD *! $ ++QKD J 7+QKD. 9 9 * D** H DE.! "#$ %! % # &&;

122 4 + G EJ! H* 9 % H &;; &;;39 P = ;++ KD>.) &;;69 H* J * = &&++KDJ 5++KD>9 + G!I!! &+ +L. DHH 9!! 9H *! JH $ J$!.( 9 $ &;;; I 9 * 9!!! H9 X E 9 D! &;;6.! I! E! J $DB $ H =c> * *! D B. ( 9 H DBE =>!* HD!!HE E$!!! J =Q> $!!* $.! "#$ %! % # &+

123 2 D 4! %! $B $! ID." E$ $* $D** HH! =!! > * H!!.( B H E D* 9 J! % $! $ H!. 4$B % =M&7! E %n& G 9 M5!9 N J J G >! E B!E! Q = 9 H &>. HH!*!I! $!H!=H &5>.!! $B 9 M5! \ *!9!! E H 7++KD =! KD9 E!* $!%!! >. ( 9 H* +++ M&7! E %!! &;;6 &;;; = H &&>. 9 ++& * HH! %* E! +++ * HH H M&7!. 9$ O %! E & G ++& %H! E + G $ $=+++>! $ $! $ ID. 4 ID B 9 +++!! ++&! =H!* >9 *! J *!* sept-95 janv-96 Truites > 15 cm mai-96 sept-96 janv-97 mai-97 sept-97 janv-98 mai-98 sept-98 janv-99 mai-99 sept-99 janv-00 mai-00 sept-00 janv-01 Amont pont Aval pont mai-01 sept sept-95 janv-96 Truites adultes > 23 cm mai-96 sept-96 janv-97 mai-97 sept-97 janv-98 mai-98 sept-98 janv-99 mai-99 sept-99 janv-00 mai-00 Amont pont Aval pont sept-00 janv-01 mai-01 sept > C &ID N +6 = N 04 =? & = & &=! "#$ %! % # &&

124 100% Différence >15 cm % Différence >23 cm % 80% 70% 60% 50% 40% 30% 20% 10% Différence (N/ha) Différence (%) % 140% 120% 100% 80% 60% 40% 20% Différence (N/ha) Différence (%) % 28/09/ /10/ /09/ /09/ /07/ /09/ /09/ % 28/09/ /10/ /09/ /09/ /07/ /09/ /09/ >>= C =D > N +6 = N 04 =? & = & &= (!9 M5! H*!I!! E $ =! &3+ &+ KD> 2 ( 9 H!!E $!* * =! > E E$!* B! %! 9 $! D!! E F $ \ HB \! 9D! E. B!E BH! * 9!I! + G H* = &+ +L >* \ E. I E H! HH E *=!!! >!!. 4$! ID 9 B J $ * % $ $D 9 X!!9! P ID $ I!. 4!H $*! D E!!! E D!! %! HE E %E H* * +G. 9 $* E D* % 9 $ * $ H!.)!I! HO9 = c> E H 9 HH J 9 *.!! E $ ID 9! D. 1 ( 4B 9 %! $ &;; &;;3." E! &;;$ D =H >9 E H C H*!* J $ 9 HE \ HB 9 E F $ = $* ^H>.! "#$ %! % # &

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

126 9 E $ F = % 9 9 > E *! J %E H D*9 HH! HH J!.) 9 9 FE$J H 9! *. EDBE! $! $!! H J. D! E H!=* c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k7l>.4$! ID!* \ 9!! 9 H* * $ 5! V! HE ID. ' $ )&9 J %! H 9!! $ E 4D HD! % =+9QKD>9E 9E $!*!!$. 4 H 97 H H* E! =739 QKD>9 * E!*! %! $S =;9L + G 59&L & G >9! "#$ %! % # &

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

128 ,-./, ,-!"#$%# 5678,-126/=.834.,- 5;63<35;-126/=.834.,- $)$* 5678, :34.,- &'(')*!*%#* "**+)* 1.0 0,5 #-#,-** 0 26/585:3,, # / =93<=<5<,;<3,8 ("'.--*! "#$ %! % # &

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

130 2! *E *! $D*S J * H * HH! = >. * H! + & D * DBE! $ * %*H. 4 * * ND* HH * + &. * * E $ $D* = H! * H ( > H * D!. " E $!*!!9 * H * $ $'=- 9&;;; ; >.4 *!* HH! * J$ 9!! 9!! $!!!. 4 H E E $'9!!! = &+! 5 K> HH =! J'*! +! 5 K>.!! H =! D $ 9! *>! H D** $'. ) 9 $ * H * $ DD 9J * H. 4!D!D* E $ D =H ++5> O * <9 $'*9 DD $'* = $'* 9!! \! % B> 9 $! D!B4B 9 * $ 4B J( D!4B E$ * $?. 4"6"0"0 / 4 -',?! $D* E!!D* E D D H! =4!%9 ++>.! $! $! E DBE $! J $!H *!! = $ > $FK." D % H!D!D* E = D D*!>9! * $!H =! 9 D $ *!E! 9J %*HH >. 4!D C! * H C!!B HH $ * 4 9DR,! * =J HH! >9!H * $! C! $D* E I $B!*! $D* E! $ H E DBE!B O $D*9 ; <..- 94!%9.9( 9. 9/.=&;;;>/* H H B!!\!9D ' 9?.B9/"19< &5&&5=&7>! "#$ %! % # &6

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

133 4 * *E * OP!! * C! "#$ %! % # &5&

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o &K+! > &7 H &79; 96 &79 +6 H N'* &39+; 9& &9 & N'* H 4B &39+; 9& &93 +# H 4B 7++=M &K+!! > &95 +9+; +9+;& H N" &9;; 9 &9+=57> &975=57> &9;; - J- +9;&, +93 )D +9 ( N'* 9&3 79;5 ( P == / 3=&K+!!!J+3+> K +7 J B +95; &93=53> +97&=53> &9 /== 2 & K +7 4B % B J / &9&5 &9&5=> +9&&=> 53K7K 9 4B * +9&; &95 /== 2= /= &++ +9&+9&7 K +7 $ 4B J( +9;66=6> &9&7 7++ & BBB% 2 $?J Y 8 * C 4 ID = Q >= F >>=!Hj * <= /&>!Hj J(< %p - q= /&3> 59 == +9;6 =&6> +9&3 =&6> && +9&&3 +=M &K+!! > ! "#$ %! % # &5

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

136 J6+L (2! $ =1-'7>H! 9 =4ID &;6>. ) E 9!* E 1-'7 Y %I! Z E $B $ *!E E 1-' =0.(D9! H 4B9!! >. 4 DBE H * $ J!!! 1-'7!! E! 1-- *! J E * B.! 4"6"4"+ $=! O * <. D $ D * $,B! &;;; $ $!!! $!!, D. 4! E E &+!!.4!! I J &+!! = >. 4H&+!!B $ %I! +=D\>J 5 9! &+J7! =? ' N' % '>. &+ 5 DBE!!.4! 5B HC H B = & J 59 ; &+>9! * = 7>9 H B E = J 6>. ) 9!D * H H E J$!=' % '>. 4 H $D*! H!! *+9&#7!hK9 *! J&9;!hK.!!!!! 9 EJH * E * $E /'. = = L = == = >=F = = KD H K! > H H. &= 8 F 4 HH %F. 4 (2 F H* *9!!! % H! E =? &? N' % '>.! "#$ %! % # &5

137 = " $ >= ( C +,, D A <A > &++ &+++ ( C 0 DI+,, & & C 4 ID ' < '? 1* 1 /J /7=&K&+!! /&>! /& 1-- /! /&=; > 1-'7 /&=; >! /= &;;6> $ A = ( 6 4 I 4 =! H >!.4 /' H* =! K&> H! E $D* H* =? N' % '>. 4 /'! * F =M+97K&%H* *>. ( 9(2 F D H* *J! &97! 5 K &!hk.)!i!! 9 9 (2! % H* *! B!E 9H *. 4$D*! H H*! *! *." E $B * (2 /'9$ H E!. ( * H D* 9 I.4 \ HB D* %I!! = >9E!!! =H Q! \ Q! H>. 4$* B $D* I J $D $ O $* 9 J D. ) 9 $! * E H!9! E$ =! > *.$!!. 4$B /'* $D* E D $ E! =' % '>. 4$ H! H *! E H! * * F! 5L +9& J &96!hK=' % '>.4!! * J * *.! "#$ %! % # &57

138 4"6"4"4 %== % " $B DE * (2=? N' % '>E * E (2 D +95!hK &9!hK E."!! +93! 5 K!!!. 4 *!! (2! J 6+L (2!%$D +9&! 5 K= F > 9! 5 K= H >.!! D +9! 5 K!!!.' E YDE Z F D!!!=+9&J7! 5 K>H J+9&! 5 K. 4!%!! (2 * E B E$J H9$*F H$ D D *E!%! (29!* H * OJE E $D*! H!. )$ >9 $B HH H!* % HH! EH* E H * < = O >. J QE =!! $!!, D > $I * E *$!=1-->!J+96! 5 K%< J &9;! 5 K.D $B!! C!)- /'!! * H %! D $!* 9 % * <! 5* HH *! % %!! 9. ) 9! $! D (2 J6+L (21--ODBE /7=^ >.4 * J +9&!hK =! *!!!> &95!hK= H >."!! +9! 5 K. 4 * H9!!!J H* *9! !5KE E $D =E E >. (? = 3)? CI+D = =DE > / +C *! ;,X (? =! 5 K> / 0 C1! E = =*!B!! 1-- =! 5 K> (2 1-- =! > 1 6+L (2 1--=! 5 K> ' ; ; < ;& &9+ +9& k+9& k+9& &+& &9;T ' 7 &97 +97; ;; +9&? & 9& &97 &7 &9; J5! D4-8-! 6 &! > 3.A+ 6, ( = > )6 CL =B== 1 = D" 4!! * /7 9! = D H > +9&! 5 K +93! 5 K9DE! "#$ %! % # &5

139 !! * 9! YD Z =+9J+93! 5 K>.! 9L &9L*! /7=793! 5 K>. ( 9 * $! 1-'7 E $ D9 H EE. 4 * *E /0) <.4! =+93! 5 K $B ; > % =! > +97&! 5 K.4 $B E *.4 * $D &+K=! H!!!9 F > 5+K H. +K!!!. */-6 =! /& % /7 (2 1-'7 6+L(2 1-'7 1 6+L (21-'7 ' < &5 ' 65; 3& +9&6? */-6 ( = )6 CL =B== 1 = D,!! * (' H % 2 C +5, #,, I <C 7, 4+, I /= 60 8 % > = > $ " % <= ! 5 K *! &9;5! 5 K>9*!! )-! K 5&+K9 D E 9!!!!! 9 H9! YD Z.!! 9 E * %< &6K &K&+!!.! "#$ %! % # &53

140 4"6"4"5 $= 9 $B E D* = /7>! *F HH$)- $ * <. HH 9$ E D* Y!B Z O9 JH *!! ^ /'. -A F = = C+;5 ID & / & = " ' E E 9 * $!!! = &;;+, D &;;;>.4 * B D %. 4! E 9!, 9!!! H9! 4D HD. $* = >D* E!* I * =E H>. *! E E DB D!E $ = F ( 49! %B %!* c>. H9 * &;;; = &3+QKD>9!! * E!!I! &;6; &;;+= 5+ +QKD>. E$HH 9 $* 9 $ HO H 9 % H!* C $! E $ % 9!! S!! = c> % (? 9 $!!* &;;69* J$* <.! &6K9E$ 7+K=&!* 5+F> &++K=& F &7!* >. ' $, 9 D* H 9 E! $ HH =Y! Z! 5 K>. 9 H* E * C &&+QKDJ$!, D 9 Y! Z QKD J $!I! 9 N J J %! /'>. 4 %E! * H! H. 4 J $!, D E $D*! E 9 $D*=/'>H J+95! H J+9$ =,B! 9++>. (9E $D* *! = /' * > H* HH H I =! > H! E =!* J >, D 9 ** * DH!.! "#$ %! % # &56

141 ) 9! $*, D = E )?!!! > I JC $! E $ = HH >9 $! E $D*DBE H $! * * J$, D. "# $ %%& '( 4$*!!!*E O $!*! $ * <. 9 * E E J $, D Y Z J *!.4 *!B! Y Z =!> $D ! 5 K9! +J65L*! %<=),9 &;;;>." H** E$J$ 9 D*! H* H E!." X! $, D 9* E! $! J H E D* EE E DBD!E O$ 9&Q!.' E $ %! 9,,! HO H *. E =HH $9 H* /'>9 HH $ * J!. 2 J * = r.&..&>! E $! * %, D! + 65L *! % <.'E J * H % < =&6K>9 * J $, D $D ;K 553K9, D! &&+K &75K.' E * )?!, D &57K9 XD 9 $ H!* 9 B.! 4"6"6"+ $= ' *!! $ * * HH! 9 D J ++!! J$ P H $J =' % >. H O $ *! * $'* =J %! $ (&& (>. 4 O *! * $'*=&96Q! > * 9 $ * H %!D* O.4 * J$*! H%J&73K=&K+!! >. ) 9 E % = D >! H! "#$ %! % # &5;

142 O 9 D $'* $'* *! =!> & &! 5 K. 4 O * $'* =Q! > I 9 %!* %*E $! *9! $!D!D*. '9!&+ *!! = % 7 = > & = = = E!>.4 E O! I!! H* H! =!*! H #H >. ( 9 * 73K9 E! &K+!!. 4! B=D $&97!>.4!!! B=D $&97!>J +!!. 4H+!!B $ %I! += >J &;+ 9! ;J7!=? 7' N' % >. 5 DBE!!.4! 5B HC H B = &J >9E = 7>9 H B H = >.) 9!D * H DD H 9 9 D E =' % >. 4 H $D*!!! *+9&#79+!hK9 *! J+953!hK. &= 8 F 4 H* HH J $!*. 4 (2 =! J&++! > 9 F H* * ++!]= > ++!]= > E H* $!* =? &7? 7 N' % >. 4 H!!! I!!E H $ B $D*. 4 E I!!! * E $D*! % F = F >. 4 /' H* = +9K&> H! E $D* H* =? 7 N' % >.! "#$ %! % # &+

143 B $ " $ >= ( C +,, D A <A > ( C 0 DI+,, & & & C 4 ID ' < '? 1 /6J /;=&K+!! /3> 1* 1 /&+J /&&=&K+!! /&+>! /&=; > 1-'7 /&=; > 1-- /&&! /&&= &;;6> $ A = ( 6 4 I 4$B /'* $D* E D $ E! 9! * F H =' % >. 4$ H! H *! E H! * * F 6 VI=' % >.4 * J * *. 4"6"6"4 %== % " $B DE * (2=? 7 N' % >E * E (2! HO H +9 &9!hK E."!! +9! 5 K!!!. 9 $ * = > I J B DE 9 E!! H E * (2 H *. 4 *!! (2! J 6+L (2!%$D +9! 5 K= F > 9! 5 K= H >.!! +93! 5 K. $B E 9! $! D (2 J6+L ( *('J +9! 5 K=! * $!> 97! 5 K= H >."!! +93! 5 K.! "#$ %! % # &&

144 = =DE > = =*!B!! (? = 3)? CI+D / +C *! ;,X (? =! 5 K> / 0 C1! E 1-- =! 5 K> (2 1-- =! > 1 6+L (2 1--=! 5 K> ' 5+; 9& < 57 & k+9& 96 ' & ; k+9&? &96 9& & ( = )++ CL =B== & %= & = D 4!! /&&=! ! 5 K> &95L 96L *! /&& =&793! 5 K>.! *! *! =79;! 5 K>9*!! )-! ;7 5+K.,!! * (' H % 2 C 0,, #6, I! C 96 47, I / = > /= 65 8 % > = > $ " 4"6"6"5 $= 9 $B E D* = /&&> H!E H 9HH $ E YDBE Z! $ /' = H!! >. J! DBE $/'9!. = =A > >>=? =A!I! $ \ E. ( 9 H! E DB*E H! H. 2 $ J $!!! =(&&(>H$*F $ &;6. =D*9 H9>9 E H* =7;QKD H9 9 QKD D*> E = >.' D* E!* H $! E $ = > %!* = HH >.! "#$ %! % # &

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hK9 *! J953!hK. &= 8 F 4 H HH %F.4 (2 F H* *9!!! %H! E =? 5? N' % >.4 =! H >!. 4 /'H* = +95K&> H! E $D* H* =? N' % >. 4 /'! * F =M+97K&%H* *>.! "#$ %! % # &5

146 " $ >= ( C +,, D A <A > &7+ ( C 0 DI+,, & & ; &+ C 4 ID ' < '? 1 /&5=M&K+!! /&5> 1* 1-- /&5 1-'7 /&3=57 >! /&5= &;;6> " $ ( +, 4 I ( 9 (2 F D H* * J! &93 &9!hK =? N' % >.)!I!! 9 9(2! H! %H* *!. 4$D*! H H*! *! *.!!!!! 9! I J D 9 B $D*.( D * J $D $ O $ $ * $. 4$B /'* $D* E! D $ E! %H* *=' % >. 4$ H! H *! E H! * * F! 5L +9 J 97!hK=' % >.4!! * J * *. 4$ *!* J *! H!!! J$. 4"6"7"4 %== % " $B DE * (2=? N' % >E * E (2D +95! 5 K= F > &95!hK= H >. "!! &! 5 K!!!. 4 *!! (2! J 6+L (2!% $D +9! 5 K =* H!!!9 F > &96! 5 K= H >.4 *!! &9&! 5 K!!!.! "#$ %! % # &

147 $B E 9! $! D (2 J6+L ( * J +9! 5 K=! H!!!9 F > &9!hK = H >.' E F = F >9 * D!!! /' = H J+9&! 5 K>.!9 * H9!!!J H* *9D &! 5 KE E $D =E E >. = =DE > = =*!B!! (? = 3)? CI+D / +C *! ;,X (? =! 5 K> / 0 C1! E 1-' =! 5 K> (2 1-' =! > 1 6+L (2 1-' =! 5 K> ' 3+& &9&& &9+ &9+ < &&+ & k+9& k+9 96 ' &&36 & k+9? & &96 &95 ; & ( = 4!! /&5=+9J&9&! 5 K> &95L 3L*! /&5=&793! 5 K>." E H9!!!J H* *9!! R HD YD Z = &! 5 K>.!! 9 E * * DD =&! 5 K>! &K+! &K+!!.,!! * (' H % 2 C 0,, + 4 I 67 8 % > = > " 4"6"7"5 $= 9 $B E D* O $!! * DD # B=3Q! >! E *! = D > H! =! H>.! "#$ %! % # &7

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

149 .!/ 4"6"#"+ $= 4 J $!! * $'* 9 O $ ("=* $'* > H $".4! E BE \ P HD! J;+@ D =' % )>.4!! E J 3+!! = >. 4H3+!!B $ 93 9 %I! &= >J & 9! &5J6!=? 3' N' % )>. &5 DBE!!.4! 5B HC H B = & J 5 &&J &5>9! * = 7>9 H B E E H = J &+>.) 9!D \ H H E J $ * E! $ H* D $ JH! H=' % )>. 4!!! $'* $! ( \ B. 4! DBE $ *!. ( 9 * J$* $'* 7++K $D J. 4 ++!! J $ * $'* E* 9E! H $ 9 * $!! 3! 5 K =!%!!> 4 H $D*! H +9& &+! 5 K * H +9& &! 5 K9 *! J &95!hK. &= 8 F > 4 H HH %F.4 (2 F H* ** J =?? 3 N' % )>.4!. 4 /' H* = +9 +9K&> H! E $D* H* =? 3) N' % )>. 4 /'! * F =M+97K&>.! "#$ %! % # &3

150 &7++ P" $ >= ( C +,, D A <A > &7+ ( C 0 DI+,, & & ; &+ C 4 ID ' < '? 1 /&J /&3=M&K+!! /&7> 1* 1-- /& 1-'7 /&3=57 > 2 +# 8 PA = ( +, 4 I ( 9 (2 F D H* *J!!hK &97! 5 K! =* D >.)!I!! 9 9(2! % H* *! B!E H *. 4$D*! H H*!! *! B!E." $B!!E H B $D*=H! 9D* 9D $B >. 4$B * (2* $D* /'* $D* E D $ E! % H* * E$H J! E *$ 9! =' % )>. 4$ H! H *! E H! * * F! 75L +9+; J &9!hK=' % )>.4! * J * *!! *H*. > 9! +9&! 5 K &! 5 KE $ +9&&+!5K9 * (2 F! =!%!!9 >. '%H* *9X H* % F.4$D*! E!! *! $HD! = >9 E F 9 $D*!%!! 3! 5 K=?? 3 N' % )>.4 $D*! "#$ %! % # &6

151 =/'> H* D** %H* * % 9 E$ H J+95K&FE$J 3! 5 K. 4$B * (2* $D* /'* $D* E D $ E!. F 9 $! * E H!J! 5 K. P" $ >= ( C +,,D A < &7++ ( C 0 DI+,, & & ; &+ ' ' 1* 1-'7 /&3=57 > C 4 ID < 1 /&J /&3=M&K+!! /&7> 1-- /& 2 +; 8 PA = ( 4 +, 4 I 9HH $! 9 H* *. 4$D*!%!!! 5 K!! *.4 $D* =!%!! +9K&> D $ E H!9!! * J! 5 K. 4"6"#"4 %== % > " $B DE * (2=? 3 N' % )>E * E (2D +9!hK E."!! &9! 5 K!!! 4 *!! (2! J 6+L (2!%$D +95! 5 K= F > &96! 5 K= H >. *! J&! 5 K!!!.! "#$ %! % # &;

152 $B E 9! $! D (2 J6+L ( * J +95 &9!hK=&! 5 K >. 4 * H9!!!J H* *9!!* E E B $D =D E E > &! 5 K &9! 5 K.! E E * D YE Z9!D H9!*. = =DE > = =*!B!! (? = 3)? CI+D / +C *! ;,X (? =! 5 K> / 0 C1! E 1-- =! 5 K> (2 1-- =! 5 > 1 6+L (2 1--=! 5 K> &9+7 &9 6& &9+ F. &3 & & +9 &&; &55; &55 +9 H & & &9 *.!%!!$ 5 &97 * F. 57 & & &93 *. & &9+ &5& +93 6; ( = & CL =B== & & <&@ =>= = &FD > " $B DE * (2=? 3 N' % )>E * E (2D &! 5 K= > 597!hK= F >." E$$*! (' 9 E * (2! E!!!!. 4 *!! (2! J 6+L (2!%$D +93! 5 K= > 9! 5 K= F >9 $* Y!%!!Z! H ('$. $B E 9! $! D (2 J6+L ( * J +93! 5 K= > &93!hK=F >."!! &97! 5 K!!!. " E DD = F > B!D D D ('!*. %! 9 F 9 (2! E$J+L (2!%9 E XH*.$ 9 *(' H J F 9! "#$ %! % # &7+

153 E$J * 9 I.) 9 $D E H * ('*. ) * 9!! $. 9 *! H* * ('!.H9!!!9 F. 4 * J! $D 597! 5 K 9! 5 K=D E >. 4"6"#"5 $= 4!! * (' H * H *. H9 /&9 * 796L 3L*! /&=&39&! 5 K>. * 9!!! +L L!I! *!.,!! * (' # (* $'* % 2 HC + 4 +A0 4 I * H C 4A6 4 5A5 4 I 69 8 % > >" = > &"!! 9 E * * $'* 7++K9 E J! &K+!!. ( 9 % $ H! D. 4! J $ * $'* H 4B U - 9 %9 $'*. ( %9! E!* H!! *E.( & 9 E $ HH HHH.4 H H* = H J&++QKD! 7+QKD > I!! =D*9 $!*!!>.4 BE =7G> %E = >! * $ = D*>! $!* E I J!!. 4 %H * $ %E * B J$ =r5...5>. E $ 9 *! * * 9! E E =E! &;;6 ++>! $ HH H =E! &;;6!B ++&>. 4$", F * EDB*E! J&3K+ &;;6 ++.! "#$ %! % # &7&

154 0! 1&2 ( 4"6";"+ $= 4$ $ H *!!D*. O $ - FE$J YD* Z.4! &++! J3! &! =*H* >97J&+!!! =* >.4 * I! H = * >9!!! HH! D %9 * $. 9 H $!! \ H. ) *! $ *. &= 8 F 4* $ H =(2> HH! * HE $ * = $ (2 *>. F 9D H +9! 5 K= H 3>. 4$D*! HO! % * H*! B!E E $D* F! J +97!hK.4 $D*=/'> % H* =k+9&k& k+95 F >9! * H* D $ =H &>.'$ 9 9! H H* * E +97! 5 K &! 5 K /' +93K&,F 9 +9K& 4D HD. D 9 $* *! H! $D* D*/' *! +9K&.! "#$ %! % # &7

155 L W 40C ) O! '. (F = C5 Z 7 K 0,,4D?F = C+7 = 0,,4D '"(C ' -. '"(C 'N -B. 40C 4 J '"(C 4 9 J -=D > -B = > L W! "#$ %! % # &75

156 200 Surface Utile (m2) pour 100 m de cours d'eau ,5 1 1,5 2 2,5 3 Débit (m 3 /s) Truite adulte Truite juvénile QMNM naturel à Pont-en-R. module réel à Pont-en-R. (37 années) QMNA5 réel à Pont-en-R. (37 années) module réel à Pont-en-R. (année 1998) Q réservé (7 l/s) 2 0, 8 $ P ( +,, = P? > == / C3D" 0,6 Valeur d'habitat (note entre 0 et 1) 0,4 0, ,5 1 1,5 2 2,5 3 Débit (m 3 /s) Truite adulte Truite juvénile Chabot Goujon Loche franche Vairon & C3)D >> F= == 3 == /"! "#$ %! % # &7

157 Surface Utile (m2) pour 100 m de cours d'eau Chabot Goujon Loche franche Vairon 0 0 0,5 1 1,5 2 2,5 3 Débit (m 3 /s) $ P ( +,, = P F= === == / C3D" 4"6";"4 %== % $B DE * (2 9 HH * 9 E H* $! E H*! *=(2 57! J77! E *! +9&! 5 KJ&! 5 K>. H* *9 D H H* =! 7+K>. F 9 $B DE * H! E * E (2D! 7+ &++K. 4 *!! (2! J 6+L (2!% 57+K= > 3+K=F >9DE $ 9 D!* E *!! B!E 9 J! =H*! >FE$J&96! 5 K. $B E 9 $ $! 1-- = E!! J +3+ K>. 4 %*! B=1-- ++K>9DE! 1-- E! -.E 9 * J $D E!!.4! $! D (2 J6+L ( * J! %!! 3+K &+K.! "#$ %! % # &77

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

159 E E! 9 $D = >. ( 9 *! H! 7+ QKD. * D** $D H $ 9!I! $H YDZ $ I *. 4$B ED* -! E $ H* 9!! H. H E HH $ $D 9! H! *! HH! D! 9 * H =!!!* %*> $D. 9 B DBD!E DB*E $D! E! =! E $ E* J* >. '! % $ 3 &2 ( 4"6"9"+ $= 4$! &++! 9 O -B,= J $! >FE$J $ E H? J;! &! 97J&+!!!.4H*! %!! H = >.4! * 9 * H FE$J * &J!!.4 H$!! H!! H J!. 9! $ H! D. &= 8 F 9!! 9 * $ (2 H * B!E \!E!%!! *$! 5 K.4! (2 *! 9!I! $ \ ++K 5++K. F 9* YD ZE.4$D*!%!! 57+!*! ! 5 K9 \ &7+ K = H 5>. 4$D*! % * H*! B!E E $D* F! J +93!hK.4 $D* %!!B = +95K& +97 F >!!E.! "#$ %! % # &73

160 Surface Utile (m2) pour 100 m de cours d'eau ,5 1 1,5 2 2,5 3 Débit (m 3 /s) Truite adulte Truite juvénile QMNM naturel à Pont-en-R. module réel à Pont-en-R. (37 années) QMNA5 réel à Pont-en-R. (37 années) module réel à Pont-en-R. (année 1998) Q réservé 2 04 $ P >= +,, = P? > ' C== 2D" E! 9 $D*!! D* =!%!! +97>9!B 4D HD =!%!! +9> *,F9 /'!%!! +93 *! &7+ 7+K=H >. 0,6 Valeur d'habitat (note entre 0 et 1) 0,4 0, ,5 1 1,5 2 2,5 3 Débit (m 3 /s) Truite adulte Truite juvénile Chabot Goujon Loche Vairon & C3)D >> F= == 3 ' C== 2D"! "#$ %! % # &76

161 Surface Utile (m2) pour 100 m de cours d'eau Chabot Goujon Loche Vairon 0 0 0,5 1 1,5 2 2,5 3 Débit (m 3 /s) $ P >= +,, = P >> F= == 3 ' C== 2D" 4"6"9"4 %== % " $B DE * H E * E (2D D 5++K 9 &7+K F. 4 *!! (2! J 6+L (2!% &+K= > ;+K=F >9DE $ 9 D!* E *!! B!E 9 J! =H*! >FE$J * =9&! 5 K>. $B E B =++K> % E E E! E$ $B $ HH. 4! $! D (2 J6+L ( * J! %!! 6+K &+K. = =DE > = =*!B!! (? = 3)? CI+D / +C *! ;,X (? =! 5 K> / 0 C1! E 1-- =! 5 K> (2 1-- =! > 1 6+L (2 1--=! 5 K> ' 7 9& +9& & < 57& ; +9& ' * *? *! "#$ %! % # &7; * 70 8 ( = ' 3 C== 2D"

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

163 &= 8 F 4$B * $ (2 H*! *!! E =D*9 4D HD >9 (2! 9 E = H9,F>$ * BE = H! YD Z>. ) D.4!%=! H4B>9H!D!D*9!! E *!!! H* HH * %!. 2!!!! E DBE H* =* J&! 5 K >.! 4&+( 4"6"++"+ $= 4$ $ H*! J$ /'. +!! (< B9 O D $ (< B. 4!! E H HH E.4!! &7+!! H %*=X >.4H&7+!!B $ %I! 5+= >J ;+ 9! J&!=? &' N' % >. & 7 DBE!!. 4! B HC H B = 7 9 & J &>9 D E = & >9E = 5 9 ;J &&>9 H B! = 3 6>.) 94B!D \ H E =' % >. )$* 9 I!!! O!$ *!. 4 H $D*! >,A64 6 VI9 *! J&93!hK. &= 8 F 4 H E * (2 $ D E =?? & N' % >.) *! DBE 9 * * (2 F.!I! 9 (2 H %H* *! * YD Z H* *. 4 H! H!.4/'H* =k+9k&> H! E $D* H* =? & N' % >. 4 /'! * F = K&%H* *>.! "#$ %! % # &&

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171 &= 8 F 4 9!! J$ D4B 9 $ Y Z * (29 E! $ H *.4 E %E JC? 9 EHE H! D E! * E E $D* H* %HH H9 E =! 9! 9 H >HE /'! $. ) 9 * I. H* HH J$!*.4 (2 9F H* * &++!]= > &6+!] = > &++! 9 E H* $!* =? ;? & N' %,>.4 H!!! %!E H D B $D*. 4 E I!!! * E $D* % F. 4 /' H* = +95K&> H! E $D* H* =? & N' %,>. = " $ >= ( A <A > 7+ ( C 0 DI+,, ++ &7+ & & &97 &97 & ' '! => 1 =+K> 1-'7 =&6 > C 4 ID <? 1* 1-- => $ B $ ( 4 4 I 4$B /'* $D* E D $ E!! * D F.! "#$ %! % # &;

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173 5!! H * $ K! DBE 9 HH!D I! ^ $! $D*DBE!! ** DE." * E!D! E! %! E 9! %." HH 9J$!! 9 H D HH H $. 4"6"+5"+ / 5.7.&.&.& 'D DBE 4 * D! J I!!D!D*!D!D*. 4 * J$!D!D*=/'>!*!!!*E =)->! JE *D O!% B= > =4B >. " $!! O * $'* 9 D=! D!B> * ( D!= $ (">4B. 9!D!D*! ^ O $! -' =? - 4B >!)- JE *D. *!! HH $ = B 4B > D! H$! E I $. 4 * H YDBE Z! DBE $D* %! * E! $ HH. 4!D!D* J! $ * DBE HH!.4?( -',? 4B! $ D! $ $ = J $ DBE H H! > HH B $! H *!!. H9 * YH Z! =!! DBE H DE! $! > H $*F % D D! J. ) 9?( $I. 5.7.&.&. 'D DBE 4$B DE *! = %J * S J $!D %! >9 *E!F =H 9!!9[>9 J$ % $!. 4 *!! "#$ %! % # &3&

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

175 4B 9 (5 D DB E ( 9 P$ HH I!9?9(5 O!!!! DB E ( (! %. 4! ** DE *!E * $ X! E!! E*E!.* *! E $ 9 B ",! F *! * * D. E $D $", 9! ^ EH9H!!!! HH H= % > $B *E E %. 9 $ HH!! 6!!!!! '?9! +! =E!! >.) 9 HH H= % >! DE. ) 9! $ *! B J $ YB! % Z! $ - \= *!! ' $ D 2 < 9 2( % 9 (!. >. B! % *! $ % HE E!B ",! '?." $ E HE H *E %.4 HE E! % * =J E!E!fE $D*! > B *E =J % > $D!!!. )! ", V++5C * DD =O 9! J >9%D $'* =O!% B>9 J'* =O 9!JH% J 4B >9 4B! * =O J DB >9 D9!!!! H D 9 %O!%. 4"6"+5"0 = = & = (BB % 5.7.&..& 'D DBE 4! ^!D!D*! *!B *E =)->! J $! * D =D $ > % D = B9H >.4 )- H $ &!hk =!>.($ E 9 D $ 9 * D * $'*! * =+97!hK>9 * D X $. 4 )- * H 9! 597 9!hK F =$ * $! )-!!!>9 *D $.! "#$ %! % # &35

176 ( * 9 DB E B=H 5+>E D $ FE$J7+! *$ $ +!hk= * &!hk>9 +! J B!!!!.* $'* 9 9 D $! &!KD $ 9J &97!K $."!!!H DBE 9 $! J!!! * $! = *!D! * * B $ >. H 9 $ 4B $! * D %$.! "#$ %! % # &3

177 7 89 9/, :,9 9 & ;6 ( &, 6 ( plage exondée hors éclusée 9-9 plage exondée hors éclusée 2 4, 8 >> P =! "#$ %! % # &37

178 ! plage exondée hors éclusée / <+, 1 +& = >=(,9<<9 > 1,<+ 2 4, \ ] 8 >> P =! "#$ %! % # &3

179 5.7.&.. 'D DBE 2!D $'* = H 5&>9! $! HE! I * = J +5!,?> $ K$ B. 4$ % )? E $ $ ++. HE! $ D =H F H >9 D % H = E!!!>. D E F 9! 9 * E =!!!!!>!*. 4 HE H!JH9 E J $!*!!9 D %=! E\!>.2! 9FE$J$! 9!* * E $ *!! F H.4 J HE * H!* 9 D % =!* ++>. 9!! 9!! * = H!*!* >. Cote de la ligne d'eau au barrage d'auberives (2002) cote (m NGF) /01/02 31/01/02 02/03/02 01/04/02 01/05/02 31/05/02 01/07/02 31/07/02 30/08/02 29/09/02 29/10/02 28/11/02 29/12/02 TRF vie sous gravier = =^ > C%2D P = C%D repro (fraie) OBR repro (fraie) vie sous gravier colonisation zones mortes = P C 0,,0 2D 5.7.&..5 'D DBD!E 4! E D!E =D!! > $D 5+&* ++59! *! % $ % = $ >. 4!DE U *! C = H >.( 9! $ ID E $! % F= $B >9 J$!!* DD =B $!* >9 J$ B=! % >9! "#$ %! % # &33

180 J $ * $'* =! %!! $'* >.( 9 $ J & &7!.)!I! HOE!! 9 $B. 4 H! D =?.5J57> * %= *.6J3&> D E. Moyenne Amplitude journalière Mois mensuelle Moyenne Max. Min Mai 10,65 2,92 4,62 0,86 Juin 13,96 4,41 7,24 2,31 Juillet** 14,07 3,55 5,27 1,45 Août 11,53 1,57 4,68 0,29 Septembre 9,56 0,70 1,73 0,29 Octobre* 8,91 0,41 1,15 0,00 7; 8 F & C_ = ` 8 & 7 <D Moyenne Amplitude journalière Mois mensuelle Moyenne Max. Min Mai 8,40 0,31 0,58 0,00 Juin 11,07 0,32 0,58 0,00 Juillet 12,50 0,28 0,58 0,00 Août 12,69 0,40 2,31 0,00 Septembre 9,74 0,27 2,30 0,00 Octobre* 8,68 0,45 1,15 0, F & $= C_ =D Moyenne Amplitude journalière Mois mensuelle Moyenne Max. Min Mai 8,87 1,38 6,06 0,00 Juin 14,07 5,46 6,42 3,46 Juillet 15,59 5,16 6,18 3,78 Août 15,84 4,75 6,48 2,01 Septembre 11,61 2,67 4,05 0,57 Octobre* 9,82 1,62 3,17 0,00 #, 8 F & = (BB% C_ =D Moyenne Amplitude journalière Mois mensuelle Moyenne Max. Min Mai** 10,90 0,96 1,44 0,57 Juin 14,47 1,35 2,06 0,87 Juillet 16,01 1,51 2,07 0,59 Août 16,22 1,57 2,91 0,89 Septembre 11,61 1,05 2,30 0,57 Octobre* 9,27 0,78 2,30 0,29 #+ 8 F & & C_ = ` D! "#$ %! % # &36

181 ! N =@> &; &3 &7 &5 && ; ( N -% - -B &! 6! &&F 7F ;F. 5F. V +V 5. &3. &. & <F A? & C 8 4, +7 = 0,,4D! N =@> &; &3 &7 &5 && ; % -B 5+K+K+5 &K+7K+5 6K+7K+5 &&K+K+5 7K+K+5 +;K+3K+5 5K+3K+5 +K+6K+5 +K+6K+5 +5K+;K+5 &3K+;K+5 +&K&+K+5 &7K&+K <F A? & $= C 8 4, +7 = 0,,4D! "#$ %! % # &3;

182 ! &; &3 &7 &5 && ; 3 7 -B - -% 5+K+K+5 &K+7K+5 6K+7K+5 &&K+K+5 7K+K+5 +;K+3K+5 5K+3K+5 +K+6K+5 +K+6K+5 +5K+;K+5 &3K+;K+5 +&K&+K+5 &7K&+K <F A? & = (BB% C 8 4, +7 = 0,,4D! N =@> &; &3 &7 &5 && ; 3 7 -B - -% &7K+7K+5 ;K+7K+5 &K+K+5 K+K+5 &+K+3K+5 K+3K+5 +3K+6K+5 &K+6K+5 +K+;K+5 &6K+;K+5 +K&+K+5 &K&+K <F A? & & C = 0,,4D! "#$ %! % # &6+

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

184 5.7.&.. 'D *E ( 9 H! E B E* % DD. ( == 4 H! $D * DD B. $!! * =&!hkj$* DD >. 4 E C J$!!* DD =!* >9 J$! DD = - >9 J$ 9 J$. ' $ % J $! DD 9 9 &;;3 &;;;! HH E JHC! BE $! $= BE!J5G>H $!!! $ 9 E! =E $!! > E! $ %E H!! J $! DD. 4! E E! J$* $'*.!! C J$!! H 4B 9 D! J$$'* 9 H J$ -. ( 9 &;;6 ++&!* E! =! *! $ HH H>9 HH!. J! BE! =!* $ > B $ =* 9** 9 F>.( J $ * $'* 9 BE!J7G9 E J H.4 * J$$'* =+97!hK> E$H* $ = &7+!> H! $. $ 9 * * $'* H $". " HH 9J 9$B HH B9 E!DE! HH C! H 9 E$ * J$$'*.4! DBE HH! 9!! (" =!%!! 3! 5 K>. ($! * % D 9J (9 H!> J $ * $'* = H 4B HH!HE %! 9 E $ %E! BE 5G 7G=H.5 53>. 9! "#$ %! % # &6

185 !H HH H 9 *! = 7L>9!! HH H = 6+L>.( * $ %E! BE 9 E $ H * J!." HH!* $! H HH H C* 9!H EDBD!E $ 9. ) =KD> &6+++ &7+++ &+++ ; ' 4 " 4? '"? /' &+K+3K++& (&3 &3K+3K&;;6 (&=+3K&&K;3> >>=> == C &ID? = & = (BB% CD ++! =QKD> &7+ & ' 4 " 4? '"? /' &+K+3K++& &3K+3K&;;6 ( (&=+3K&&K;3> 2 4# 8 == CGID? = & = (BB% CD! "#$ %! % # &65

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

187 +" $>>= DD & 5 7 D & 5 7 '* & 5 7 4B & 5 7 4B ( D & 5 7 4B ' & 5 7 0" %= DD & 5 7 D & 5 7 '* & 5 7 4B & 5 7 4B ( D & 5 7 4B ' & 5 7 4" = > DD & 5 7 D & 5 7 '* & 5 7 4B & 5 7 4B ( D & 5 7 4B ' & 5 7 5" )? DD & 5 7 D & 5 7 '* & 5 7 4B & 5 7 4B ( D & 5 7 4B ' & 5 7 6" = DD & 5 7 D & 5 7 '* & 5 7 4B & 5 7 4B ( D & 5 7 4B ' & 5 7 7" = F DD & 5 7 D & 5 7 '* & 5 7 4B & 5 7 4B ( D & 5 7 4B ' & 5 7 #" $? a DD? -B?* D? -B?* '*? -B?* 4B? -B?* 4B ( D? -B?* 4B '? -B?* ;" 2 a DD? -B?* D? -B?* '*? -B?* 4B? -B?* 4B ( D? -B?* 4B '? -B?* 9" ) => DD? -B?* D? -B?* '*? -B?* 4B? -B?* 4B ( D? -B?* 4B '? -B?* 2 4; 8 = = = '- F? B $= F C=F D! "#$ %! % # &67

188 > DD & 5 7 D & 5 7 '* & 5 7 4B & 5 7 4B ( D & 5 7 4B ' & 5 7 0" P * 4". DD & 5 7 D & 5 7 '* & 5 7 4B & 5 7 4B ( D & 5 7 4B ' & 5 7 5". = DD & 5 7 D & 5 7 '* & 5 7 4B & 5 7 4B ( D & 5 7 4B ' & 5 7 6" = =? CD DD 1 & 5 7 D 1 & 5 7 '* 1 & 5 7 4B 1 & 5 7 4B ( D 1 & 5 7 4B ' 1 & 5 7 DD ( & 5 7 D ( & 5 7 '* ( & 5 7 4B ( & 5 7 4B ( D ( & 5 7 4B ' ( & 5 7 DD /N & 5 7 D /N & 5 7 '* /N & 5 7 4B /N & 5 7 4B ( D /N & 5 7 4B ' /N & 5 7 DD <N & 5 7 D <N & 5 7 '* <N & 5 7 4B <N & 5 7 4B ( D <N & 5 7 4B ' <N & 5 7 '- DD & 5 7 D & 5 7 '* & 5 7 4B & 5 7 4B ( D & 5 7 4B ' & = = = '- F? B = C=F >D! "#$ %! % # &6

189 4 HH!DB!E! %! J.4D $D** % * *!!!. 4 B H! J DD HD!!. 4! E * DE H & E H! J $ E =! %! >.! 9 DE E = $ E(1>. 4!fE $D* H 9 * 9! %."!* E DBE % E *!fe $D* H $D!.4! $E HH D!#. 4$B HH H DE % DE J $ HH *E H.! %!E E!# 9 HE $E E *. 4$ J HE H 9 E! $ *DE =' > 9! $!*. E E *E 9 D! $ E J! E! ='* >!* E E! J *!.! %!E 9 H HH H= 1> % D 9 E D! ",9 E BH! = DE $D*> % D $.) 9 D %!E 9 E $E* E! % D E! %! J.)$! 9 E DB*E! ++5 * H! J!. & & 8 & & &A >> < C= ( % D = & > == =B= & A =A == &= >= C= =D" A = C= = ( %A = > D! "#$ %! % # &63

190 4"6"+5"4 = = & = 5.7.&.5.& 'D DBE 4! ^!D!D*=D>9!D!D*9! *!!*E =)->! J $! * D =D $ > 4B 9 $ % ' =!D*>9! D!B =D>9 4B =D>. *!! 9$*! HOH* )-E 9 J$ $ 4B =E F 9 $'* >. ) O9 )- H! +9 &9!hK! = H > $ =! $! >.( $ E 9 D $ 9 $ % * J$* E! $ 9 H 9 * $ 9 E$!!.4 )- * H * 9! 7! 5 K$ =!>9 *D 4B $. (4B 9 * =! 5 K> H!! =9! 5 K>.4! =&+! +J! 5 K>." E!!! 9)?! * $ J9! 5 K *!! D =+! >. H* * $ 9 D $! 9!! J$!! H!D.4 *! * E H!. D!!**! J$ H! * O DB E. *! E H DBE * H EE.) 9D! H= * H H 9 >9!**! 9!!* \ HB. 5.7.&.5. 'D DBE ' 4B 9 E H! 9! H!. HH 9 H Y Z C E $ HH 9! *!hkue $ H J 7595!,?9 $I = * * >.4E * J! 5 K9 H H $ 9 *!. 4E *!5K9 H =& J D F!!!>9 O! % $!! *. ( $ ++ =? >9 H F9!.4 F H! H* * H $ =V!* >9 * H H J DKF."! 9 D % =!*!* %! >9 H F E.! "#$ %! % # &66

191 )$! 9 * H! $. * $HH HO * D E B EE = 9! F >9 E H $!*!!9 4B =J$ $ 4B >. Durée journalière de fonctionnement des groupes de l'usine de Bouvante (2002) durée (h) = =^ > C%2D P = C%D TRF vie sous gravier repro (fraie) OBR repro (fraie) vie sous gravier colonisation zones mortes 2 5, 8 >= = C 0,,0 2D 5.7.&.5.5 'D *E ( == (4B 9 H! J$ H 4 J $!!! 9!! =5>9! H % J$!! 9 J J $ $ = > D!*B =7> F!* $ 4B.4 H!! $D!! *! = H & >." E H &;;9 H J $!! = 5 >.! "#$ %! % # &6;

192 ) =KD> +++ &7++ & /' '"? 4? (5 ( ( >>=> == C &ID = & = CD = > +995! =QKD> /' '"? 4? (5 ( ( == CGID = & = CD = > +995! E!H! 4B 9!! J $!!. 4 E* = H &;;> J %!! $! H! = >9!H E $!! X =!! >.' E * $D 9!! $* $ J? B! $! *D!!9 E %E!.4 D**! E E $ * E$!*! * E H! 9! H! * DE. ( 79 HH H!!I! * H. *! $! 9 E!H! "#$ %! % # &;+

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

194 4B 9 E!! = > E! 5!!. 4$B DB*E J $ (B! % $ J $ HE = &.+#H.++>!! * =J$ 6!E > E*E =J $ 7 B > DE D =H FJ >. )$! 9!E E! $ * DE E!# 9! *! 4B. ) 4B 9 $!DB!E E E* J$ $ 4B E$! * %!. 4 D $D** % * E E F!B 4B.4B H! 4B. 4! E * DE H & E H! J $ E =! %! >.! 9 DE E = $ E(1>. 4!fE $D* H 9 * 9! % 94B =!! >.4$D $D* H 4B $ 4B E$!!B!."!* E DBE % E *!fe $D* H $D!.4! $E HH D!#DE $. 4$B HH H DE % DE J $ HH *E H. 4 HE E E *!# 4B 9 E!E $E. 4$ J HE H 9 E! $ *DE =' > 9!$!* J$ 4B. 4 *E E! 4B 9!=E* > CE!!B J* D9! $ 4B.! %!E 9 H* HH H= 1>4B D9E D! ",9E BH! = DE $D*> % D $. 9 D %!E = (>9 $ =/$> $ $E* =<$> 4B =! % $D*>. )$! 9 E DB*E! ++5 4B H! J! 9! ++59 $ 4B = O D!! > E I * =H* c>j$* $.! "#$ %! % # &;

195 4"6"+5"5 = = = 5.7.&..& 'D DBE * $!! 4B 9 D. HH 9 * D %$ 9 $ 4B $! H = &J7 % $ >. 4 4B!*! J!!! J $ * $'* =! *D >. 5.7.&.. 'D *E E!! J$! * * DE I * J$!! 4B J'* U$! $!*. )$! 9 * H * $", J '* 9 E $", D %!.) 9$B!! J % HH!. 4"6"+5"6 = & = 2 C= D ( $ 9 H J$! )?=! )75&>! $ $' 4E J(. 4! H! $D!! = HH H *!? >. 4!! % 9E $H! 9!!?.( %9$D $! $ HH H9E! *!. 4 D*!!9!! %. 4$H!! HH J! * C D 9 E $ 9[ 4$!! E $D? $".! "#$ %! % # &;5

196 ! =QKD> 7 + &7 & /' "? ( H >>=> == C &ID? = & = C2D < +994! =QKD> 7 + &7 & /' "? ( H == CGID? = & = C2D < +994? 4"6"+6"+ / 2! $ E DBE $D*!D! )7(=!DE F ++5>.!D! $ E DBE $ J$ EH EHC * 9D H9*!DE =!! "#$ %! % # &;

197 HH >. E %OC * DD J B9 4B * $ 4B J( D! B=O!J >. ) %!! $ E $D*DBE * < $'* 9E HD* % =HD D % >. 4"6"+6"0 % ( 9! * DD - OD! C! $ * DD! DD = &>9! DD J = >9 - = 5>. (4B 9 E OE H EH$*F $ B EDBE % EE.4! E O C? B J $ FE$ )76=4B &>9 )76FE$ )3+=4B >9 )3+FE$ $'9=7! D >J$ )3=4B 5>9 $'FE$ $ 4B =4B >. 4! O $! * DB!DE = > $. ( 4B! F B! HD* = $'9 $ 4B >. 4$ I H * YH ZY*H Z EDBE!! * E. H9 $* H B! H 9 $ I J $ = * >. DE! H 7 'JU ='> HJ H!!! J!.! "#$ %! % # &;7

198 ) = $= * ( C*(D /&&& ';+ &5+ ( #+JG+ j=/g'>%% ' M7+ ' M7 ' M7 (! G&+KG+ ' M ;7 WE* &+KG&+ 57++# ; W 7K&+ &7++# ) &6 ) && 5 ) &55? +K7 ) ++#&7++ k& k&& k& (! HH *9 $D k++ #5 8 = & 4 D E! =D9 9 9 *>E EDBE * * H. L ) CI +++D = CI 9,D $= CI +4,D 5,D T * = P 40 & & &97 & &+97 & & ; & &;96 &9++ &&; 2 & & &9+ 7&9+ 9+ &&97 & &93 9; &97 5+5& #6 8 %=> >> = * = P C LD & 57++ ( EDBE & &7++ & % = >> L! "#$ %! % # &;

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

200 P = 8 4 J N * N'* 9 B * H. 4! * &! D. 4 D D DE * N 5+!. 2 B! D! N! *! * J! * N =! N J N >. B! N H!. 4N! =!> H J N * E E! R s! s *. 2 H =* >!! *. ( + 0 C * D % $= C '! 9!* H N! sdbe se N * J N H! E N!!. 4N =J N> * s* s E! N % J N * =D >. 9!*!! * J HD J! *!. ( 4 C ( 5C N 4 H9!! N!*!!9! N E J! HB I *!! E (!!!. ) 9 H 9 D N! N *! E!! H! = % N N!>. 4!* *!!9 E HB J N! =D! sd!s>9 N!!! E. 4 J N H N HH J N N B! J =!>!E!. " I B *E =!* 9 > J N * 9 HB B &! "#$ %! % # &;6

201 )$* && 9 $!* 9 J$$ $I H H!!."!* E JD! $ E J$ 9* HH $ H!.4$ I $ %! % H$.) 9 *! $ 4B 9 $'9 *!E.4$$ HD! = J> D D = 5 7!>9E! E! $ $$. ' E$ % 9 E E \ HB % 9!!! *H* J HE$J$!*!!. 5 = $ 0, 8 = 5"+ ' &> = 4, $%/BE )D$)?!!!!!!! DBE =! (HH (V /BE ((/>.! E E *$ 9 $ 9 * 9$!!! ((/U! E %. $ +++9$H!,/$)? C * 5+++ HHD =! 9 9!> 9 $!*D &! H! *9! * %E J 9 $ $ )J$ )!!! SD $ 9 &5 $H!* % 9 F J( ( 9 $ E? D D!)9! %$ $ OJ$ *!- %. && * =(,'/#,'0>! "#$ %! % # &;;

202 @A A< &9 (, 9 & : ( < 2 5# 8 =! "#$ %! % # ++

203 ! F 9,/ )D $)?!! $H! =HHD 9! 9! > O =!!!! %! >. )!I!! 9 )?9 E? )! ID - 'EE ID $! %E! = H J $ 9D I H %>E E ID *. -!! $H! 9!* E E E ID a* 9 K t J ID *!! = H!!!E ' ' ID - 'EE >.$ 9 P$ 9 H E ID 9 * * =)?> K ''-'C = *>$)?E! $ $9 * ''-' H! D E H! DBE P ID $. )$ 9 ID!! * = DD $'* %! >9! *$)? ''-'9 % ID =H 57>. * ''-' X $ HE =H* 9!B! >9 %E I! ID =! $ %! >9! =9 $D > HH $.' E!!E 9 H! BD *.! "#$ %! % # +&

204 ''-' B ((/ $ P ''-',! L! N'* N'* - H= C+D C0D & = > & = >! 9! =* ND >! 9! ( 5 DD DD #!! ( DD!! DD 5 ( & D* D* 5 ( &!! & 1 = 5 ( & B> N'* 5 GD & '* #H 4B H 4B - - (\ #7 8 (= H=! & 5 D & &! 5 5 5"0 $ = = ( * ''-'9 * ID C H 9 $!! K Y Z 9 H9! $ $. 9 $''-' B E E E J $! ID = *! $'*PID 9! * $'* DD >. 4$''-' 4,! $ HH!! * $D* DBE! V % H* HE (\ BJ $HH. $ HH B! =**J +!hk $ #**D U D D>. D! 9 $! $9 E I *% C B9 E D $ +!9 * $'* 9 $ E!!! * = &+!$ I * >. H * 9 \!!! * $ * E 9P* = P! "#$ %! % # +

205 $! E $ >9 * $'* =!!! $'* 9 D 9 -!!!! H 4B >9 $ E D FE$J 7+! = H >. ) 9 9 E * D =H!%! H J7+!9 H J&!K>9 * E $ =H &! K & &97!K>.E 9!I! $ H 9 =ID %> E $ E a t FE$JH $ 9-9 H 4B =!! J >9 $!! = * $ 4B >. ( 4B 9 E * E $ 5+!.4! %$ HH &+! E % H.- 9! $ $ P *$ H* D $. 5"4 4 *! E I = 9 D*! \ * >BDOJ $ &;;6(,'/",'0!. ( / 9 H! * H B =!!! 4 %! >. $* \ * =! 9 %, B!9 \! H! 9 ( D >. 4 D! <! I -. 9!-J$!! I D $! = $&! $ >.)!I!! 9 H! DS = D! DD > * E E $! DD = > H*."!!! 9! D E *S! % H!!! D % D!$. ) 9!!9 *S! $ B % % D* $ R J B \ *. 4! - % (\ B! D! $. ( HH 9 \ *! C H! $ *S!? 9! "#$ %! % # +5

206 D* H = Q Q H H > 9 9)&7=J HD!! >!? - 9 D!! = E E D* > O 9 /4-8? =P!! >. ' $! * - 9! = DS >9 \ Q ))$' \ =FE$% %- - >9 9 *. "!!! 9 \ * ' -.) O9 E H!,9 D*! H! 9 D* 9 D!, -9! DS u D* H J$ ) &+9 \ $!. - 9 D! $ =++ J 7+! >9 *! * E E D*! \ * = J >.' 9 D* D H! *.) 9DBE < HH % EE D*.) 9 D!$!!D. H9E E D*%,9%, E$ D* D!, *. )! * 9 \ * H =FE$J7+! (' D* %! >9! F %! H! D!. 9 J $ O! =D! % B 9! 4 > H! D* 9 H! *. H BD 9*S! $ *D $D* J $! E \ *.,, 9 D!$!% * E * - $ ID -! -.' %!9 * H )? B % D* J H =H. >.! HH 9 H $! BD * D! $ % $ % D! % I.BD 9 *! E!F H * $ %HD!! )&+5!!!.($'9 *! $ 9$D* *! )&+5*. )4B! DB E 9 E $.!! 9! D= D9, $'P! *!! $ 9 E $ D*>.4!, *! *.4 *! 4B $ Q )) D J (< B X!*!% $.9$ d' - D*J D!*B! *D!$.4 D 9 HH 4B 9 *.( *!! "#$ %! % # +

207 !% E *E. $H! HH *? =! $ (,'/",'0! > E 9 $ HH.! "#$ %! % # +7

208 /'" 5C2(',(? "() 4$'2 )(-"4"2:'12' "12( 4 %! %EE $ C DB 9! * 9! 9 9ID 9!.4! $ B J$* HD ' % 6. $ DE BD HD C % $ 9 9 B 9 $ 9 H! *F H9 E! F. 4$ HH D! D F $! F %E.! "#$ %! % # 0#

209 + == +"+ =? 4! DB E N H J6+++ Qi9! J!!." % &7++.!* 9 H DE 397! 8iD9 &97 L. % )?9!!* N! N 9 4!! 7++Qi! J= J H $*F $ >H!! J &.&+.&;&;$DB J $ &;;. +"0 4!!. 4! U H H J. +"4 =! D! U J 9E %!. 4$ J )?!!. 2 * BD E!. =C! $'* $ * H$*F $ HE 9 E $!! * * K K!!!. +"5 == & 4/! JHH %C B!!!! = >E.4 H 9!! E C!! DE! E % *! J = O!! 9!B H J7++!>. H * $ H!! $ 9 E! H *E E $.!! "#$ %! % # 0

210 EH $!EE *! H R * = H!* $ &+!!!!!!! >9 H9!H!! E * J * 9! HD*. ' $ 9 E$ $ $! $ * 9E* J! \J HH. +"6 > <=> 4 *F H E! I H! % C ' C! H E DB $ H$.(! P F %!! $ H* 9H!!* I $'=-.? 9$('2"!! (- $ EJ$ $ >U H I!!I!. ' $! C DB E H! H! %EE 9E$!!!%!!! E$*.4! J C o o o E * %*! = E H > D U!! % * $ R U E J H! HD! H * O. H9 E E *F H * 9 DB J C H$! I D D! %9 H!!%!!! E H X.4$ % * HH J I.! "#$ %! % # 0,

211 ## 8 F = == ==! "#$ %! % # 0

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