Parc Naturel Régional du Vercors Adresse postale : Chemin Fusillés 38250 LANS EN VERCORS Tél. 04.76.94.38.26 OUVRAGES HYDRAULIQUES : optimisation de la gestion des régimes de débits pour les bassins versants de la Bourne et du Furon (affluents de l Isère) OPERATIONS B4-2 ET B4-6 DU CONTRAT DE RIVIERE Tome I - RAPPORT D ETAT DES LIEUX DES MILIEUX ET DES USAGES Adresse Postale : 62, Boulevard Niels Bohr B.P. 2132 69603 Villeurbanne Cedex Tél. 04.78.93.68.90 Fax. 04.78.94.11.98 Portable : 06.70.93.44.55 Email : contact@asconit.com G2C Environnement Adresse Postale : Parc d activité du Point Rencontre 13770 VENELLES Tél. 04.42.54.00.68 Fax : 04.42.54.06.78 Email : g2c@g2c.fr La Bourne en amont du Pont Rouillard Décembre 2006
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4 \ D!!E* 9H! 9!* H! =* >. % %$ C = %E >!B d"?? B & B H I DB*E 9 DE D E 9, $ B &7 8! $ 9D 9 / =! 4> \ D! d"?? B &.( &3D $ E ID E 9D E DE 9 B $ = H 4B > 4B (< B9 d"?? B &9 %!!! $I!! +++ C D* D % = H H! H9 D*> J D 9 H = H 9 % 9D*9* >9 *! 9! 4B = % > ( =J $ %!?>9!! D 9 HH!,, 9E HH! * DD 9! D! QE!E*.! "#$ %! % # 5
- 56&3 F!E* =A =A = - > = 56 ( DS 563 N B 56! +& + 56+5 56+ 56+7 56+ 56+3 '+3 " * " * " * " * " * " * " * d "! % +++&++ d"?? B & +++++ d"?? B & +++3++ d"?? B & 56 ++&&++ d"?? B & 56 % -D! 4-4 56 4 * * % DS 6+ +++! -/ +;K+;K&;9 * ( +97DJH* B!E S ( 7DJ B!B! *!9 S F* ( 93DJ B!B D! *! % R 56 4 *? ( 796DJ B!B 56 DD H DD =, > ( &5 D J H B =E H!E* > 56 4 * ( 9&DJ B!B 56 O * DR ' D!9(' 9 % 4D 9(' D 9D 9 B9( B D 9D 9(< 9(- / % HI!!* D 9 9 9( = % E >!B N' D +++++ d"?? B & 4-4 ( 7 D J B!B =HI! E D > & D & D D = 9 < 9' B9? 9 B 9,N [> 5;& D 9 HI9 J I DE 9 *E 9!E HE =D!9 9 H[> 6 D N D! J I DE 9 DB*E 9 K!D* = 9! 9 9 H! > ;6 D N J IDE 9 DB*E 9!!!H =* H* J JH! > &+33 D HI9 * J I DE 9 *E 9!E 9E 36D!* D! JI DE 9*E 9 K!D*! "#$ %! % #
- +53+ d"?? B & +53+ d"?? B & =A =A = - > = 9(< B9( - 9(< 19 % +53+3 d"?? B & 94 9!*\ &&3+7 d"?? B & &&3&+ d"?? B & 56 7++++ d"?? B & 56 (< B9(4 B 94 9 ( 9(\ -D (< B9(4 B9(\ B9( D! B9 ( B9 '* B9 B9(< %94 -?F?I 4 ( -?N2 I,!* 9 4B!* 4, N B B 4B ;;++++ d"?? B & 4D!* N 56+3+ d"?? B & 56 56+3+7 d"?? B & 56 (- 9 4 4D 9(' 9(- 56+3+3 d"?? B & 56 (' N/ *B N'*?I D %* -H 56+3+6 d"?? B & 56 4D ( B 567++++ d"?? B & 56 9( 9(\ -D ;7 D HI9 * J I DE 9 *E 9!E!!!H =7 9 H H > 565 D J I DE 9 *E 9!!!H 9 K!D* =D!9\ > &5 D J I DE 9!!!H 9 K!D* =!H9 H9 > 655DJIDE 9E 9B. (! = *!* DR ' > 7&+ D D 9 9 9! D %JIDB*E B.&78! 9 D 9 *D 4 N I DE HE ;;D N JIDE 9 DB*E 9B.4 B B! E. D 9 H D* 9!*!! N'DR D 9 9!E9 HD J I DE 9*E 9 K!D* = R)R! > 3&D JI*E =JD 9 e 9 > 7D JI*E = C % 1! 6 " 6 31" > &;D HIJI*E!E =! 9 > 6+ D HI J I DE = > 5 D HI9 * J I *E 9 DE 9!!!E 9 DE =D 9 H > 5635++++ d"?? B & 56 9(' B ( 75D HI9*JI*E =D >! "#$ %! % # 7
- =A =A = - > = 563++++ d"?? B & 56 '- d D! N'- 5636++++ d"?? B & 56 '99( 563;++++ d"?? B & 56 D 9DD 9 9 5667++++ d"?? B & 56 4 9 4 5666++++ d"?? B & 56 +5 d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fE ND* N = H H! H9 %! 9 D*9 * 9* > &6;+D QE =B%9**9* >! "#$ %! % #
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+"; / =!! % $ C E! =!! 4 FE$ HH U *, >9 E! = O 9)D 9 F D 9 4B!4 >9 = 9 - E! F J$!>9 =!!D!* 4> = ' 49 9 % B U,, U $9 B9 9 (? HH B>9 4 &;5+ B 9 9 $ 9H H 4B $ %!?. ' F =E &;;7>$I * ='> $!* 9 JH $".! "#$ %! % # 6
0 =? = 0"+ = 4 55!! R!!! *! %B. " C!!!! J$" 9 ) 49 )R! C!!!! 4 BB9!!!!. " %! 6!! B!! $ $! C ("-B9 ("' 9 (-' #?I 4 #?$2 9 (-' 4B 9 -H 9 9 J $" 9 E B B =! HD! >. 0"0 $ 5 8 = = F 3= ( +999 +9;0 +999 4 &;;; $ 57++ D*!! C6L BB#! 9L B 1-9 6L BBR!.4 *! B 1 -! R!. )$ $ * -9!! E!! J $ = X *! $! $ >9!B H* \ D* =&+D*8!] 9&7J+D*8!] B>.! &;6 &;;;9! \!! =DS 9 9 (- (\ -D >!!! 9 F$D&+++ D* =4 9 P % $! 9 - U % &>. 9!!! &;6 &;;; =57L J (- >. E 9 *!! $D ='9 O 9-4 / 9 % / >.! "#$ %! % # ;
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= O > - = - >. 7+++J&++++ J'9 J $! - 9 &+ +++ J 7+ +++ J 4 =!! P! +++ 7+++D*D>. 4! $!!H * $!!* E! = 9 9 E9[>. 4! J $ I * HH C gqbq9h DB E 9 DD JH $".2 H $ J( 9! J %! 4 9 <! B9 O =J $ O >9 - = ' - >9 H BD 9 H % % %!?9 B E!! DD?9 $. ( $*-= &;66J&;;>9 E %J * -! * I J.4 =&;;3J++>9 9 E! H! D!!* * - =!* &;;39!* &;;69 F9 V!* &;;;9 F!* +++9 F ++&9 V!* ++> 6&5H%! *. 4 =? $! J$ % 4B = (< B> = H >E!. ( H $*F H $!! * $ 9!! H 4ID 94$9 ()', ).44ID 9!* $ $ 9! H% *." I! D* $D! )!! EE, =),>.! "#$ %! % # 5
4"+ H= C09 < +9;5D = 540B6 $ & 44YID Z@67&;F&;6E!! J$ 9 C &K $ " 540B6 $! E H =&K+! *!B =! > $ &;69&K&+! &;69&K+! % &;6H % CD9DR 9! M6+! 5 K E *!! I * &K+! >. 4!!H $ 9 *!!*E I! &K&+!! $ FJ. $! J6+!hK9 $ H% *!!J! H E &K+!!. K $ " 540B7 $! E H! *. ( $ 9!! I E $ H *. %9 $E $H!* 7 J * =*! > $ $! 9 5K $ " 540B9 $! & & @ $. 4"0 $= +6 = +990 ( 9 * *! $ *!!!! = H J $ 4 57 $! >.! "#$ %! % # 55
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4"6 %F C= 0 +7 = +9+9D 4 4 +7 = +9+9 = >!H $ 7 4 @6+75& &7F &;6+. H!! C =7++Qi>9 H = $! EI *E >9 $9 Y Z E DBE = H% ) $>. 4"7 ' == 4$ 4555 $! E $ % $ ID! * 9 X!! $*! Y Z. 2 Y)!! EE, Z=),> *?)! ID 9 * ( ID." Y(D! )! Z = ()>! 7 *E E!! $! 9 *! J!. 4"# = & '? C'D 4$5F &;;9 ()',! H EE! % H.! $ E! $!! * H *F H E E $.) 9 *F H H%! E* *E 9!!! * $. 4 H * $H!.! )*-! E =)-> DBE. 4 )- *!!! 9 EE = ()',4 >.) 9 % H! $BI H ()',. ( * DR - 9 * *F H % H * E *F H E J!! JC H! DBE! * $ HH 9!!D %DBE! % % 9 $ = D*> H **E!!! E $! "#$ %! % # 57
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!! 9!! $ *. 2 @ @ 3BB / @ / 3 @ (BB % P @ B /BB 3= @ @ B1BB $? +976 +977 +97# +97; +979 +9#, +9#+ +9#0 +9#4 +9#5 +9#6 +9#7 +9## +9#; +9#9 +9;, +9;+ +9;0 +9;4 +9;5 +9;6 +9;7 +9;# +9;; +9;9 +99, +99+ +990 +994 +995 +996 +997 +99# +99; +999 0,,, 0,,+ 0,,0 0,,4 ) ) ) +7 8! "#$ %! % # 75
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Débit mensuel d'étiage de la Bourne aux Jarrands (m3/s) 1,4 1,2 1 débit en m 3 /s 0,8 0,6 0,4 0,2 0 1949 1947 1950 1961 1962 1964 1952 1966 1957 1959 1955 1967 1970 1971 1985 1986 1969 1978 1953 1965 1976 1998 1973 1958 1963 1983 1980 1997 1984 1979 1982 1974 1948 1960 1981 1956 1972 1987 2001 2000 1996 1975 1954 1951 1968 1995 1977 1999 2002 ( &;;7++5 ( &;3&;63 Débit moyen de la Bourne aux Jarrands (m3/s) débit en m 3 /s 4,5 4 3,5 3 2,5 2 1,5 1 0,5 0 1964 1947 1957 1962 1969 1967 1986 1961 1997 1952 1976 1985 1998 1954 1968 1979 1960 1975 1951 1987 1980 1978 1977 2002 1995 2 + &? 1! "#$ %! % # 73
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X * OS %B!! $)? )"=! *E /B>C *^+ &;3J++=! F J$!* <>9 *O&& &;;7J++=*J$! DD J $ %$)?!! >9 *O& &;3+J++=S J! - >.,S J 9 %E! C = => = =5> B! $E9 X! HH OC!* $ % U Y Z J **D. 4. 0"+"0"4 F & 4! O! E %9J!! $)? (". 4!! )? $DB C $ J *. C %* <9! $'*9! H YH $ Z $H Q $. D * 9 C = + + 4 ** * D $.4 * E! %<$X *!!. % $, D 9! * C H$ % J$D *! E $ &;&;.X **! "#$ %! % # &
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4$!* * %7 C 1 / 1 1 B = B = 1, D =! 5 K> &9& &9; +93+ &957 1, =! 5 K> 9; 976 &9+ &9;5 HH &33L &55L &;L &5L 06 8 F @ ' - 1 DE =! 5 K>?! 1" 2" / / 1 1" K " =" -" =" / 9;+ 9+5 957 593 59+ 9+; +9+ +93 9++ 9;5 9; &96 9 &933T 1, &933T 1, &933T 1, &955T 1, &955T 1, &955T 1, &9;T 1, &9;T 1, &9;T1, &95T 1, &95T 1, &95T1, K 07 8 %= @ ' - P +99;.&...5 O, D,, D, 9! 9 HHU * &+Q! 9 $ J\ $!,. 2!! *,'0! $ $! D, D 9 * *HE, D.! "#$ %! % # 7
!!!! = *! 9 HH %*! >9! O$ &;;6C 1 / B 1 1 B = B = 1, D =! 5 K> &9& &9; +93+ &957 'O5=! 5 K> +96 +9; +9& +9 HH +L &7L &3L &L 0# 8 F L ' = B ' - 1 lo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
4 $'*9 9!!* J*!B!hK * +!hk. 4 D$'*9! $'* 9! E! D! *!=+!hk>. 4 = C&6!>9! 9!! E E E H 9! *! 6+!hK.! '. S J! * D. 4 I!! \ B $'* $!. 9 H! J!! D $ %! * $'*. 4 4 9 J&;8! $'* 9!! $ E*!B +9!hK.4 H 9 D $ *$ 7+!hK= 4 >! D $* ++!. 4 E! 9 * *9!! $! \ B.( $! =<.<., 4! 9?D '&;36>9 \ B %*D C! % $'* % = 4 >. 4 4 H!!! D! $E* 9 %! 9 9! % $'* =D$'*>. ( % % % *! $ *9 $!!.)D 9 H! ( (< P % H!*! H. '$D 9!* E $'* E 9 E!H HO C * $! H $'*9! H* 9 % $'* H U * % J H * H9 H 9 $'* ** * J E U!! '*9H %. = & E $'*!!!H 9 $ %! *9 $ E E F $* E $.'9/. H! * $'* * J$! < $ &;5. H H! E J $ &;;6 H* $'*=!B 97! 5 K>9!* $! =$E$X! DBD B!$* >.! "#$ %! % # 3
) * * *DBE!! * 9 * * $'*!! $E=>* D. * H C = 4 *! $'* $ &;;6 * C = / 1" 2" /" / 1 1" K " =" -" =" & 1=! 5 K> 97 79+ 7955 &9+ 796; 9;; 9&5 &9;6 9;+ 393 39+ 797 79;5 4, 8 3? = & P +99; 4 * *!! HE *!! C %! YZ ( $'* 9 E % $!I! U $H. = 4! 9H9!! $D9! % % QE 9 E %!I! %I!!! %. "!*! $! = * >9 = * $DB9!!, D %! >. 4 * $ % $)? E % $!, $ $! $ *. 4 D $ * FE$J D 6+!C! %!! * H! 1j!4D D=!j+9567 H!H )?>. (! ++& ++I!J C E * 9! * J H E$! I =H >. C! 9 HH! %* 9 J! % $! 9! QE! % %.! "#$ %! % # 6
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$ % =' % ;>. D D $* H*!E = HH H> =*! >. 4"5"0"+ 4 * * B ( ID $' 3. ID E = 9 9ID > &;65 ++&. H9 $ * =!9!* H! > I! HH H *! = H >. $ B E! H!9JH$ = >!= >. ) F * 9 J 6 9J+ 9&+ * 4B 93 * 9 -?. 4 E *. ($!* * =6>9.!! 9 =&;;+> 9H H $! = J$ * DD ' >. 3 9,. B!9<..=++>(BD &59 %HH. ( ID '&69.;G %.&;97! "#$ %! % # ;3
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
$7 + (! = 4>9Y!Z ' E &;;5 ++5.! * $! J $H DB E. E DBE HH = $ \ >. I!! H. 4 H D* % 9 E! ** E H*B %. % 9 4 HH H *%!! *! = 6+++KD9H 7>!I! %I! \HH =!!! 7+++KD &;;!%!! &7+++KD +++>. ) 9 F $ 93L=&;;;>J9L=&;;3>!B $ +L $ =H >.4 + G $D 7+=+ G >KDJ ;+++=+ G >KD %!I!. $ I *9\ F %. Effectifs de 0+/Effectif total (%) 70 60 50 40 30 20 10 0 Truite fario (0+) - Station Pont des Asniers Densité Proportion 10000 9000 8000 7000 6000 5000 4000 3000 2000 1000 0 Effectifs (N/ha) 22/09/1993 28/09/1994 1995 03/07/1996 25/06/1997 17/06/1998 11/06/1999 18/07/2000 11/07/2001 09/07/2002 27/06/2003 2 5 8 < & C, R D > ( = 8 $ (H= " E J $! + G H*. HH 9 B D =!*!I! >! E!!! %!. H $!* * 9 % H D &;;5=* >. %=& G > = G > DHH D E 7L 5+L.! "#$ %! % # &++
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
!! 9 H* HH H + G * &;;;! E H* 9*!=7 3+! 5 K H!>E$J$! = &5+! 5 K * >."$ * H! H* 9H E $ ++E = F %!* >.! F=769! 5 K! 3F%<9579! 5 K DD > %E H* = > + G *! F!I!. 4$ HH! 9!! $B H + G! E$ DBE E $* ^H! E $! =9&;;79)H 9 ++9.9 ++5>. 4 *!! *!!I! E HH H!!!* &7QKD=++>!%!! 5+QKD &;;5= ++ QKD +++>!B $ 7+QKD. H = HH H *! >!* % H $. " E 9 SD \HE 9 9 %E.$!! $ +++E *!! 9EI D SD E H E E! J $ $ =- >." E$ $ * H! E.) 9 9 $H %. H9$! ID B!! $ J.' E &;; +J5!$!*! $".! "#$ %! % # &+
Biomasse (kg/ha) 500 450 400 350 300 250 200 150 100 50 0 Truite fario - Station Pont des Asniers Effectifs Biomasse 16000 14000 12000 10000 8000 6000 4000 2000 0 Effectifs estimés (N/ha) 22/09/1993 28/09/1994 1995 03/07/1996 25/06/1997 17/06/1998 11/06/1999 18/07/2000 11/07/2001 09/07/2002 27/06/2003 Biomasse (kg/ha) 500 450 400 350 300 250 200 150 100 50 0 Truite fario - Station Camping de la Balme Effectifs Biomasse 16000 14000 12000 10000 8000 6000 4000 2000 0 Effectifs estimés (N/ha) 1993 1994 1995 10/06/1996 25/06/1997 19/06/1998 1999 2000 2001 2002 2003 Biomasse (kg/ha) 500 450 400 350 300 250 200 150 100 50 0 Truite fario - Station Colombier Effectifs Biomasse 16000 14000 12000 10000 8000 6000 4000 2000 0 Effectifs estimés (N/ha) 1993 1994 1995 1996 04/11/1997 1998 12/10/1999 2000 2001 2002 2003 Truite fario - Station Passerelle Béaudet Biomasse (kg/ha) 500 450 400 350 300 250 200 150 100 50 0 Effectifs Biomasse 16000 14000 12000 10000 8000 6000 4000 2000 0 Effectifs estimés (N/ha) 1993 1994 1995 1996 05/11/1997 1998 11/10/1999 2000 2001 2002 26/06/2003 Biomasse (kg/ha) 500 450 400 350 300 250 200 150 100 50 0 Truite fario - Station aval Lyonne Effectifs Biomasse 16000 14000 12000 10000 8000 6000 4000 2000 0 Effectifs estimés (N/ha) 1993 1994 1995 1996 1997 17/07/1998 1999 2000 10/07/2001 2002 25/06/2003 Biomasse (kg/ha) 500 450 400 350 300 250 200 150 100 50 0 1993 1994 1995 Truite fario - Station Pont de Manne 1996 1997 Effectifs Biomasse? 6 8 > C>>=> D 7 B = 8 $ (H= 17/07/1998 1999 2000 10/07/2001 2002 25/06/2003 16000 14000 12000 10000 8000 6000 4000 2000 0 Effectifs estimés (N/ha)! "#$ %! % # &+5
4$B * '! E HH H %!* 9 9!! $B 9 * H* J5!.( =&;;5++5>9 HH HH HO!. * \ M+! &;; &;;9E$! +++9 J D! HH E E!. 4 HH H %!! ++& ++5.- 9 $!* 9!* J* Densité d'individus (N/ha) 900 750 600 450 300 150 0 Truite fario - Station Pont des Asniers >20 cm >23 cm juin-93 déc-93 juin-94 déc-94 juin-95 déc-95 juin-96 déc-96 juin-97 déc-97 juin-98 déc-98 juin-99 déc-99 juin-00 déc-00 juin-01 déc-01 juin-02 déc-02 juin-03 2 7 8 > @ = C0, = <& +995A 04 = = D ( = 8 $ (H= 4$ HH ID HH J! DE D!! &;;= +J5!> E$ * 9 HH $.!9! I D $!* $! + 5!. 4 JH. Proportion (%)/ population totale 16 14 12 10 8 6 4 2 0 Nb individus/ha Proportion 800 700 600 500 400 300 200 100 0 Nombre d'individus /ha 22/09/93 28/09/94 17/06/95 03/07/96 19/06/97 17/06/98 11/06/99 18/07/00 11/07/2001 09/07/2002 27/06/2003 2 # 8 CID 0, 04 = C XD @ "! "#$ %! % # &+
"! E! J $ &;; P D &L9! 7L!E."!I! B $9 E H!!!!! &;;9 ++ ++KD9H +++= SD c>.9 *!! E E HH $! E '. 2" D*9 HH HH HO! 9 H * *! HH H &;; &;;=E $ 9$ $ 9H 6>.4$* $ &;;7! D *.4! 9! HO! 9 &;; &;;;!* *! E 9F! * &;;5 &;;. ( *!! ++5! J * &;;5. 4 D* * E % E $ $D*! *!. ( H! BH!! = 9 E H $ J $!9[>.)!I! HO9 J$E* =!J!* J * *>! H!. 2 ) 9 ' 49 H!9E 9J 9 H! $ H!! DB E E$ $B F HH. 4 * ' H 9 FE$J 5+QKD =&;;5>9!!* I H!!H! E! H! *!.!9B!E!* HO!!! H $DB = > $! HH H + G. S9 *!! = H 7>!I! I =!! c>h. HH 9 E! 9! 9 E! H B!E.!H!! H $ R E ID E.( % =! > B *F H $!!!DE =!! >! $. 4 *!!! HH ID 9 $! $ * *.! "#$ %! % # &+7
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
( $!* 9 X E* 9! H E* F $ =+ G > HH H! D &;;3.4 H* + G * * 9 E! &;;;. $-= +1 9 ) 9 = >9 H &;6+ $.' E HH $9 H 9$* D*! BH! B! =*! E $ % F $ 4>. ' 9!-J! = % * $! 9 J / *! >9 H D* = &;; &;;6>.44D HD $ E$ &;; &;;3 E!.! ** = E H *B >9! *. 4 HH H H +++ KD *! H J7+QKD=H 7>.) HO \ 9 + G H* &;; &;;6= &L> &;;3= 6+Lm>. DHH C * HH H +G9 H* &;; &;;6 = 5+++GKD> H &;;3= 5++++GKD>9 H* S. 4 H &;;3 $ %E!* %! = * $! *! 7!>." E + G!* J * ' 9 HH *! = &J&+!! &;; &;;39 $J &J! ' >. 4$B D!! HE! E!! E Y!Z9 $! $ HXD. E $!= > % 9 H*! %! =+ G J& G >! H* * = H J 7!! F>! D!! E! D K*!. 4! H 5+L=D &;;3> &37L=D &;;>9DHH E H$! $S &;; &;;3. % =& G J G >9 %! 5+ 7+L. " * E J $ 9!!!!E H*! + G = HH H!> *! F $ H ID. ( 9 D* * = > - J $ < =, D >. 4! "#$ %! % # &+3
! J $! E $ 9 H!* QE =, D 9 >.4 HH H! %I!! H* =! 5++KD>!I! E *! =H J3QKD>."!* B *! &;; =$P HH H!E >9D EY!! Z FE$ &;;3=*! >!E!* * H. 2 4 /!H! E DBD!E $ 9!! * D! * =E *>."!* E!! %HH * < =, D 9!! >9 * *! E $ 9 E!* H! B DB*E! * =",9 >. H9 * $ * < = F ( 4> E! J$ E 9*. 4 H! E D* H! H.)$ (9 $D** H* = H > E * E!.) 9!9!E = J!* D HD* >!! $ D!. H9 $ * =9 9 H *>9!I! 9!! *! E E E. HH 9 = J $E* > $ B!! H*.! "#$ %! % # &+6
Biomasse (kg/ha) 90 80 70 60 50 40 30 20 10 0 Chabot - Station Pont des Asniers Effectifs Biomasse 6000 5000 4000 3000 2000 1000 0 Effectifs estimés (N/ha) 22/09/1993 28/09/1994 1995 03/07/1996 25/06/1997 17/06/1998 11/06/1999 18/07/2000 11/07/2001 09/07/2002 27/06/2003 Chabot - Station Camping de la Balme Biomasse (kg/ha) 90 80 70 60 50 40 30 20 10 0 Effectifs Biomasse 6000 5000 4000 3000 2000 1000 0 Effectifs estimés (N/ha) 1993 1994 1995 10/06/1996 25/06/1997 19/06/1998 1999 2000 2001 2002 2003 Biomasse (kg/ha) 90 80 70 60 50 40 30 20 10 0 Chabot - Station Colombier Effectifs Biomasse 6000 5000 4000 3000 2000 1000 0 Effectifs estimés (N/ha) 1993 1994 1995 1996 04/11/1997 1998 12/10/1999 2000 2001 2002 2003 Biomasse (kg/ha) 90 80 70 60 50 40 30 20 10 0 Chabot - Station Passerelle Béaudet Effectifs Biomasse 6000 5000 4000 3000 2000 1000 0 Effectifs estimés (N/ha) 1993 1994 1995 1996 05/11/1997 1998 11/10/1999 2000 2001 2002 26/06/2003 Biomasse (kg/ha) 90 80 70 60 50 40 30 20 10 0 Chabot - Station aval Lyonne Effectifs Biomasse 6000 5000 4000 3000 2000 1000 0 Effectifs capturés (N/ha) 1993 1994 1995 1996 1997 1998 1999 2000 10/07/2001 2002 25/06/2003 Biomasse (kg/ha) 90 80 70 60 50 40 30 20 10 0 1993 Chabot - Station aval Pont de Manne 1994 1995 1996 Effectifs Biomasse 1997 17/07/1998 2 ; 8 $ C>>=> D 7 = 8 $ (H= 1999 2000 10/07/2001 2002 25/06/2003 6000 5000 4000 3000 2000 1000 0 Effectifs capturés (N/ha)! "#$ %! % # &+;
$ & ' 7 1) 2"" * * DD - 9 $ \. $2 ( J $!! * DD =& Q!9 $>! * *!! 9!I! 4D HD * E = HH H *! >H* =! 7QKD>. 4 HH H!! &;;3 &;;; =H >9! J* HH H + G = + G 369;LJ9L>.! * F $ I %E * DD =H &;;;> K % H * E O!! =H! * &;;;>. * * H ' E + G %I!! H* &;;;9!! HH!. ' $ 9!I! 9 *!!! =G&6QKD>9! $ J $!! * =D &;;>9 E!* % D &;;3.' E J ' 9 *! *! H =;+QKD &;;3 7 QKD &;;;>. " D %!! 9 ' 9!! H!. % BE 9 * I!*!I!. 4$* &;;6! $! D % H.!9$* &;;6 +++! $ HH $S 9*! E." HH EJ$! * DD H 9E!!E!* E.4$! D %H $I! =,B.9+++>. 4 ID B!! =*!* >9 ** E!$ HH H + G H* 9!! * J = H! >.) 9 $B $HH HH E! 9&Q!J$!9 * HD* DD. ) H9 D!! + G E! *! = 9&;6;>9 E H $! $S 9!. D 9 HH H D*!!I!! &;;3 &;;;C*! D*! + 6+QKD HH H $! 5+++ KD =&;;3> J 7+++ KD &;;; =H 7>.!! J!* % D &;;69$ * I =* &;;6> E$H =! J$!!* DD H! H* >. 4$* &;;6!.! "#$ %! % # &&+
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
2 '$!!* DD 9!!! % 9J D* H.4 $ HXD! * DD H H*! D*E!!. 4!* H* J!!! J. 4 *! J! =H ++ QKD>! I *! '. HH 9 $ HO E 9 % BE E % *! 9*!!! $! $ = E $ F HH H9! $ * $! F $ >. )$ 9 &;;; H$! % *= * DD > * $. 4 D % H $ I H 9!! $* $ &;;6 +++9! '!*! E!! D F $.4 ++5= >! E *! H* E$ &;;3 &;;;9! $*! D!! = ++> E J!! * &;;;9 B! B J$E* \H*!! $* $HH * \ H K! B!. H9 $ $! DD H EE! H9 % $ H H* = K! (29D!D*>!. 4 = HH HK! (2> *. (! -!* <&;;6.&;;3.&;;;.&;;3 &;;;! =K! (2> 9 +97 39 &39+ 697 +9+ +95 &9+5+ &9;6+ &9&+ HH H=*K! (2> F +9&&6 +9+& +9 +95;; +9&; +957 +93&6 +9+3 &9+5; +9&& 5# 8 CI 0 ( D CI 0 ( D & C( D" = 8 ' ="A 0,,," $* &;;69 HH! 9 $! DE $S =!! D &;;6>$.! "#$ %! % # &&
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
2 ( J $ * $'* 9! X E & 9 E $ HH H HH.) 9!! %* 9 HH*."$ H $ O E I! =D*9 H9!*!!>. ( 9 BE =7 G > %E! 9! * 9!!!! H* =! >."!I! D* E H*.!9 $ H! %E H* 9 $!*!!9 E I J!!. 4 * %!! DB E E C * $'* E * 7++ K *!=FE$J3!5K>! 9 B E H 9 $ E!! * $'* 9 J 9 $ 9* J= > J % H 9 D 9 %B D!. J $ *! $ H! =D > E! H J * H BD! E $ =%B 9 /9[>. 4 H*! $ H DHH!. 4 H $!*!!! B H H*. 4 * *E! $!! = > J $!*. 9 *! 4B D! H*.) 9 $!* $ DD 9 E %! 9 E! * J *=* >. 4 H! B = > = HH H 4B FE$J H >!! 9 F 9 * 9! *!=#!>J!! P! HE. % D! ID $ E H!!! ID.! 9$! %,!9$ J! $* H* HH H E 9 HH $! $!.)$ HH H! E 9 B!*!. 4 D! 9 E %E H * $!*!!* &;;6 ++&9 $D! &;;+9 H* J $!*!! E $! E! $ +@." 9 $HH*9 * J! *.! "#$ %! % # &&7
(!!DE =!* %! > H! H* E J $! HH * E $ %!."!! H D!E! * $'* (\ B9H E$ 9!* D! =H > E $!*. 5...5. 4 - ( $ 9E D &;6 &;;6. (! =! >9!! &;;3 9J 9 H 4D HD.' $* D*9 EX!! E $ E!* F! B. 4 D*! * &;6;&;;+ % $ H - = <, D >. $ E 9,.4!* I J*! E E $. 4 9!! ++ QKD H HH H!! F $ =;L + G >.! H! E!9 $ $* D* $. = J $!,9 N J J $!! - >9! DJE &;; &;;6."! E 9 J 9$D J &;;=+9QKD>.H9 9* $ &;; *!! &;;=396QKD> &;;3=&597QKD> * &;;6= &57QKD>. Nombre d'individus/ha 6000 4000 2000 0 1992 1993 1994 1995 1996 1997 1998 CHE TRF Nombre d'individus/ha 100 000 80 000 60 000 40 000 20 000 0 1992 1993 1994 1995 1996 1997 1998 LOF VAI 2 9 8 C>>=>ID = @ & ' C/D C%2D $ C$)D CDA 3 C3D = >= C2D CD" '$ 9! H!.4 &+QKD &;;J! &QKD &;;69 *! 4D 9 +QKD=&;;>J &+QKD. $! E$ &;;9!B = &++8KD> * J! 7+ 5++8KD &;; &;;39 * H! J 659QKD &;;6. 9 &;;&;;39!!* J * '!. &;;3 &;;6E 9 *! $! E HH H!! "#$ %! % # &&
!*.! H $ $ B! $ H* % &;;3 &;;6. 4$B D!!! &;;6 H* * $!* $S F $ =+ G >E E 795L =5;9KD>.)$ HO 9 + G H* E$ 79&L=&;;>J795L=&;;6>9 E! *!." $ * H!!! HC E! * $ =FKF >9 J E =k3!> HH J J $ 9 E $!! F $! *! 9 E \ HB E E! H* =! >. 4$ E E 9 $ %! D &;;9 + G! &;; $ 3 KD E $ 9!I! $S &7&79KD. H9 9!! *! * &;;6. E 5+ QKD =&;;> 73QKD=&;;3> D *! % 7+ QKD &;;6.($ $!* E J H$D =H* HH %$! %! >9 H E! 9!! $! D 9 *E 9 E D E. E! J E $ % F!! -. I! %! $ E! H! * -. * E 9! $K9 DHH!! $ 9 $! E! $ H* $!* $. 4$!! $ H* D 9! *! HD! H* D*. 9J%! H 9 %$ $ < &7+!9D &;;3.( %9!! J H9 4D HD!! HH. HH 9!9 =&&6 QKD> 4D HD =&5 QKD>! E H*! =7QKD>.'$9$ $ E J =&3QKD>94D!! H* 9! &97 QKD &97 QKD.!H $ %E E $ E$ $ % F % =! &7+!>." $ E $ HH E K E $D* DBE.(!9 E H* % =4D HD 9>9E 9 E H* J =! * 9!* % *9 H9.>" $ J % E HH E DBE % J D! $ HH!**! E % H. D! B! =J $!!! >9 $ % E Y! Z H.! "#$ %! % # &&3
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
Biomasse (kg/ha) 500 450 400 350 300 250 200 150 100 50 0 Truite fario - Station amont Pont d'echevis Effectifs Biomasse 16000 14000 12000 10000 8000 6000 4000 2000 0 Effectifs estimés (N/ha) Biomasse (kg/ha) 500 450 400 350 300 250 200 150 100 50 0 Truite fario - Station aval Pont d'echevis Effectifs Biomasse 16000 14000 12000 10000 8000 6000 4000 2000 0 Effectifs estimés (N/ha) 28/09/1995 01/10/1996 23/09/1997 22/09/1998 1999 2000 21/09/2001 28/09/1995 09/10/1996 23/09/1997 22/09/1998 07/07/1999 22/09/2000 21/09/2001 2 +, 8 > C >>=>D = @ & CD @ & CD &=" Effectifs de 0+/Effectif total (%) 80% 70% 60% 50% 40% 30% 20% 10% Truite fario 0+ - Station amont pont des Echevis Densité Proportion 8000 7000 6000 5000 4000 3000 2000 1000 Effectifs /ha Effectifs de 0+/Effectif total (%) 80% 70% 60% 50% 40% 30% 20% 10% Truite fario 0+ - Station aval pont des Echevis Densité Proportion 10000 9000 8000 7000 6000 5000 4000 3000 2000 1000 Effectifs /ha 0% 28/09/1995 01/10/1996 23/09/1997 22/09/1998 07/07/1999 22/09/2000 21/09/2001 0 0% 28/09/1995 01/10/1996 23/09/1997 22/09/1998 07/07/1999 22/09/2000 21/09/2001 0 2 ++ 8 < & C, R D > 3 &= 4 HH H + G!! * =H &&>.4 $D! 7+KD=&;;79&;;69++&>J 6++KD =&;;3> &7 3+L=&;; &;;3>.4 Y! Z9! H*! * E 9 * E ' =!>. 4 H* &;;6! = $ E E *! E >. $! 3"! '$$D 9 E!!* J!C H. 4! * = D*> H* = 4D HD >.' E$ ++&9 HH H H* E % E 4D HD!* HH=55+KD>. H9 HH H B!E!!E!C $D +++KDJ &7+++ KD *! $ ++QKD J 7+QKD. 9 9 * D** H DE.! "#$ %! % # &&;
4 + G EJ! H* 9 % H &;; &;;39 P = ;++ KD>.) &;;69 H* J * = &&++KDJ 5++KD>9 + G!I!! &+ +L. DHH 9!! 9H *! JH $ J$!.( 9 $ &;;; I 9 * 9!!! H9 X E 9 D! &;;6.! I! E! J $DB $ H =c> * *! D B. ( 9 H DBE =>!* HD!!HE E$!!! J =Q> $!!* $.! "#$ %! % # &+
2 D 4! %! $B $! ID." E$ $* $D** HH! =!! > * H!!.( B H E D* 9 J! % $! $ H!. 4$B % =M&7! E %n& G 9 M5!9 N J J G >! E B!E! Q = 9 H &>. HH!*!I! $!H!=H &5>.!! $B 9 M5! \ *!9!! E H 7++KD =! ++ ++ KD9 E!* $!%!! >. ( 9 H* +++ M&7! E %!! &;;6 &;;; = H &&>. 9 ++& * HH! %* E! +++ * HH H M&7!. 9$ O %! E & G ++& %H! E + G $ $=+++>! $ $! $ ID. 4 ID B 9 +++!! ++&! =H!* >9 *! J *!*. 4000 3500 3000 2500 2000 1500 1000 500 0 sept-95 janv-96 Truites > 15 cm mai-96 sept-96 janv-97 mai-97 sept-97 janv-98 mai-98 sept-98 janv-99 mai-99 sept-99 janv-00 mai-00 sept-00 janv-01 Amont pont Aval pont mai-01 sept-01 700 600 500 400 300 200 100 0 sept-95 janv-96 Truites adultes > 23 cm mai-96 sept-96 janv-97 mai-97 sept-97 janv-98 mai-98 sept-98 janv-99 mai-99 sept-99 janv-00 mai-00 Amont pont Aval pont sept-00 janv-01 mai-01 sept-01 2 +0 8 > C &ID N +6 = N 04 =? & = & &=! "#$ %! % # &&
100% Différence >15 cm 1400 180% Différence >23 cm 300 90% 80% 70% 60% 50% 40% 30% 20% 10% Différence (N/ha) Différence (%) 1200 1000 800 600 400 200 160% 140% 120% 100% 80% 60% 40% 20% Différence (N/ha) Différence (%) 250 200 150 100 50 0% 28/09/1995 01/10/1996 23/09/1997 22/09/1998 07/07/1999 22/09/2000 21/09/2001 0 0% 28/09/1995 01/10/1996 23/09/1997 22/09/1998 07/07/1999 22/09/2000 21/09/2001 0 2 +4 8 >>= C =D > N +6 = N 04 =? & = & &= (!9 M5! H*!I!! E $ =! &3+ &+ KD> 2 ( 9 H!!E $!* * =! > E E$!* B! %! 9 $! D!! E F $ \ HB \! 9D! E. B!E BH! * 9!I! + G H* = &+ +L >* \ E. I E H! HH E *=!!! >!!. 4$! ID 9 B J $ * % $ $D 9 X!!9! P ID $ I!. 4!H $*! D E!!! E D!! %! HE E %E H* * +G. 9 $* E D* % 9 $ * $ H!.)!I! HO9 = c> E H 9 HH J 9 *.!! E $ ID 9! D. 1 ( 4B 9 %! $ &;; &;;3." E! &;;$ D =H >9 E H C H*!* J $ 9 HE \ HB 9 E F $ = $* ^H>.! "#$ %! % # &
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
9 E $ F = % 9 9 > E *! J %E H D*9 HH! HH J!.) 9 9 FE$J H 9! *. EDBE! $! $!! H J. D! E H!=* c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k7l>.4$! ID!* \ 9!! 9 H* * $ 5! V! HE ID. ' $ )&9 J %! H 9!! $ E 4D HD! % =+9QKD>9E 9E $!*!!$. 4 H 97 H H* E! =739 QKD>9 * E!*! %! $S =;9L + G 59&L & G >9! "#$ %! % # &
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
,-./,-012-34.,-!"#$%# 5678,-126/=.834.,- 5;63<35;-126/=.834.,- $)$* 5678,- 93585:34.,- &'(')*!*%#* "**+)* 1.0 0,5 #-#,-** 0 26/585:3,, # / =93<=<5<,;<3,8 ("'.--*! "#$ %! % # &
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
2! *E *! $D*S J * H * HH! = >. * H! + & D * DBE! $ * %*H. 4 * * ND* HH * + &. * * E $ $D* = H! * H ( > H * D!. " E $!*!!9 * H * $ $'=- 9&;;; ; >.4 *!* HH! * J$ 9!! 9!! $!!!. 4 H E E $'9!!! = &+! 5 K> HH =! J'*! +! 5 K>.!! H =! D $ 9! *>! H D** $'. ) 9 $ * H * $ DD 9J * H. 4!D!D* E $ D =H ++5> O * <9 $'*9 DD $'* = $'* 9!! \! % B> 9 $! D!B4B 9 * $ 4B J( D!4B E$ * $?. 4"6"0"0 / 4 -',?! $D* E!!D* E D D H! =4!%9 ++>.! $! $! E DBE $! J $!H *!! = $ > $FK." D % H!D!D* E = D D*!>9! * $!H =! 9 D $ *!E! 9J %*HH >. 4!D C! * H C!!B HH $ * 4 9DR,! * =J HH! >9!H * $! C! $D* E I $B!*! $D* E! $ H E DBE!B O $D*9 ; <..- 94!%9.9( 9. 9/.=&;;;>/* H H B!!\!9D ' 9?.B9/"19< +++9.79.9.&5&&5=&7>! "#$ %! % # &6
*E!C!!! ID.!D 9 H $ $ %!E!D!D*.!D*!! E $D* = + &> >= = $D*% H! >9 H *9 HH K =! >! $ =! > E % H $! = 9 D 9! * >.'9 9!!.4 D H! *. 4 J! &++ D $ *! E &7 O $ H &7 J 5+ H $ J %* HH * H * *.! E!%!!DJ DE * *! J! ^!D*E $B.!D O!! DBE H E *! *E = D DDE > C $ -! -B U $!? - 4B =.'>4B. ) 9! DD 9P!D*!! ^. 9 *! %! HH!!D* \."!*! E!D %E HH $ $!D*. 4"6"0"4 ( &!* E!D!D*! ^ 9$!* DE $.4 E! D H * $ (2 H! *E O.! $ H % =' % >C ",9 D E! Y$!* Z 9 H!! DBE 9 * DBE 9 *! DE EH E 9 H DBE!! B %! DBE 9 $/*=/'> 9 F 9 H H * H **. * $ (H 2 =(2> * $D* 9F 9 H H * H 9! "#$ %! % # &;
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$B E 9! $! D (2 J6+L (21--.4 * J +95 &9!hK=&! 5 K >. 4 * H9!!!J H* *9!!* E E B $D =D E E > &! 5 K &9! 5 K.! E E * D YE Z9!D H9!*. = =DE > = =*!B!! (? = 3)? CI+D / +C *! ;,X (? =! 5 K> / 0 C1! E 1-- =! 5 K> (2 1-- =! 5 > 1 6+L (2 1--=! 5 K>. 65 59+ +953 &9+7 &9 6& &9+ F. &3 &9 +95 +95& +9 &&;+ +95. &55; 9+ +9+ +9+ +9 &55 +9 H 37 96 +9& &933 9+ 9 77 &9 *.!%!!$ 5 &97 * F. 57 &+96 +96 9 597 56& &93 *. &5 9+ +9+ +96 &9+ &5& +93 6; ( = & CL =B== & & <&@ =>= = &FD > " $B DE * (2=? 3 N' % )>E * E (2D &! 5 K= > 597!hK= F >." E$$*! (' 9 E * (2! E!!!!. 4 *!! (2! J 6+L (2!%$D +93! 5 K= > 9! 5 K= F >9 $* Y!%!!Z! H ('$. $B E 9! $! D (2 J6+L (21--.4 * J +93! 5 K= > &93!hK=F >."!! &97! 5 K!!!. " E DD = F > B!D D D ('!*. %! 9 F 9 (2! E$J+L (2!%9 E XH*.$ 9 *(' H J F 9! "#$ %! % # &7+
E$J * 9 I.) 9 $D E H * ('*. ) * 9!! $. 9 *! H* * ('!.H9!!!9 F. 4 * J! $D 597! 5 K 9! 5 K=D E >. 4"6"#"5 $= 4!! * (' H * H *. H9 /&9 * 796L 3L*! /&=&39&! 5 K>. * 9!!! +L L!I! *!.,!! * (' # (* $'* % 2 HC + 4 I @ +A0 4 I * H C 4A6 4 I @ 5A5 4 I 69 8 % > >" = > &"!! 9 E * * $'* 7++K9 E J! &K+!!. ( 9 % $ H! D. 4! J $ * $'* H 4B U - 9 %9 $'*. ( %9! E!* H!! *E.( & 9 E $ HH HHH.4 H H* = H J&++QKD! 7+QKD > I!! =D*9 $!*!!>.4 BE =7G> %E = >! * $ = D*>! $!* E I J!!. 4 %H * $ %E * B J$ =r5...5>. E $ 9 *! * * 9! E E =E! &;;6 ++>! $ HH H =E! &;;6!B ++&>. 4$", F * EDB*E! J&3K+ &;;6 ++.! "#$ %! % # &7&
0! 1&2 ( 4"6";"+ $= 4$ $ H *!!D*. O $ - FE$J YD* Z.4! &++! J3! &! =*H* >97J&+!!! =* >.4 * I! H = * >9!!! HH! D %9 * $. 9 H $!! \ H. ) *! $ *. 4"6";"0 @ &= 8 F 4* $ H =(2> HH! * HE $ * = $ (2 *>. F 9D H +9! 5 K= H 3>. 4$D*! HO! % * H*! B!E E $D* F! J +97!hK.4 $D*=/'> % H* =k+9&k& k+95 F >9! * H* D $ =H &>.'$ 9 9! H H* * E +97! 5 K &! 5 K /' +93K&,F 9 +9K& 4D HD. D 9 $* *! H! $D* D*/' *! +9K&.! "#$ %! % # &7
L W 40C ) O! '. (F = C5 Z 7 K 0,,4D?F = C+7 = 0,,4D '"(C ' -. '"(C 'N -B. 40C 4 J '"(C 4 9 J -=D > -B = > 2 +9 8 L W! "#$ %! % # &75
200 Surface Utile (m2) pour 100 m de cours d'eau 180 160 140 120 100 80 60 40 20 0 0 0,5 1 1,5 2 2,5 3 Débit (m 3 /s) Truite adulte Truite juvénile QMNM naturel à Pont-en-R. module réel à Pont-en-R. (37 années) QMNA5 réel à Pont-en-R. (37 années) module réel à Pont-en-R. (année 1998) Q réservé (7 l/s) 2 0, 8 $ P ( +,, = P? > == / C3D" 0,6 Valeur d'habitat (note entre 0 et 1) 0,4 0,2 0 0 0,5 1 1,5 2 2,5 3 Débit (m 3 /s) Truite adulte Truite juvénile Chabot Goujon Loche franche Vairon 2 0+ 8 3 & C3)D >> F= == 3 == /"! "#$ %! % # &7
Surface Utile (m2) pour 100 m de cours d'eau 600 500 400 300 200 100 Chabot Goujon Loche franche Vairon 0 0 0,5 1 1,5 2 2,5 3 Débit (m 3 /s) 2 00 8 $ P ( +,, = P F= === == / C3D" 4"6";"4 %== % $B DE * (2 9 HH * 9 E H* $! E H*! *=(2 57! J77! E *! +9&! 5 KJ&! 5 K>. H* *9 D H H* =! 7+K>. F 9 $B DE * H! E * E (2D! 7+ &++K. 4 *!! (2! J 6+L (2!% 57+K= > 3+K=F >9DE $ 9 D!* E *!! B!E 9 J! =H*! >FE$J&96! 5 K. $B E 9 $ $! 1-- = E!! J +3+ K>. 4 %*! B=1-- ++K>9DE! 1-- E! -.E 9 * J $D E!!.4! $! D (2 J6+L (21--.4 * J! %!! 3+K &+K.! "#$ %! % # &77
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
E E! 9 $D = >. ( 9 *! H! 7+ QKD. * D** $D H $ 9!I! $H YDZ $ I *. 4$B ED* -! E $ H* 9!! H. H E HH $ $D 9! H! *! HH! D! 9 * H =!!!* %*> $D. 9 B DBD!E DB*E $D! E! =! E $ E* J* >. '! % $ 3 &2 ( 4"6"9"+ $= 4$! &++! 9 O -B,= J $! >FE$J $ E H?. 4 597J;! &! 97J&+!!!.4H*! %!! H = >.4! * 9 * H FE$J * &J!!.4 H$!! H!! H J!. 9! $ H! D. 4"6"9"0 @ &= 8 F 9!! 9 * $ (2 H * B!E \!E!%!! *$! 5 K.4! (2 *! 9!I! $ \ ++K 5++K. F 9* YD ZE.4$D*!%!! 57+!*! +9 +93! 5 K9 \ &7+ K = H 5>. 4$D*! % * H*! B!E E $D* F! J +93!hK.4 $D* %!!B = +95K& +97 F >!!E.! "#$ %! % # &73
Surface Utile (m2) pour 100 m de cours d'eau 400 350 300 250 200 150 100 50 0 0 0,5 1 1,5 2 2,5 3 Débit (m 3 /s) Truite adulte Truite juvénile QMNM naturel à Pont-en-R. module réel à Pont-en-R. (37 années) QMNA5 réel à Pont-en-R. (37 années) module réel à Pont-en-R. (année 1998) Q réservé 2 04 $ P >= +,, = P? > ' C== 2D" E! 9 $D*!! D* =!%!! +97>9!B 4D HD =!%!! +9> *,F9 /'!%!! +93 *! &7+ 7+K=H >. 0,6 Valeur d'habitat (note entre 0 et 1) 0,4 0,2 0 0 0,5 1 1,5 2 2,5 3 Débit (m 3 /s) Truite adulte Truite juvénile Chabot Goujon Loche Vairon 2 05 8 3 & C3)D >> F= == 3 ' C== 2D"! "#$ %! % # &76
Surface Utile (m2) pour 100 m de cours d'eau 550 500 450 400 350 300 250 200 150 100 50 Chabot Goujon Loche Vairon 0 0 0,5 1 1,5 2 2,5 3 Débit (m 3 /s) 2 06 8 $ P >= +,, = P >> F= == 3 ' C== 2D" 4"6"9"4 %== % " $B DE * H E * E (2D D 5++K 9 &7+K F. 4 *!! (2! J 6+L (2!% &+K= > ;+K=F >9DE $ 9 D!* E *!! B!E 9 J! =H*! >FE$J * =9&! 5 K>. $B E 9 1-- B =++K> % E E E! E$ $B $ HH. 4! $! D (2 J6+L (21--.4 * J! %!! 6+K &+K. = =DE > = =*!B!! (? = 3)? CI+D / +C *! ;,X (? =! 5 K> / 0 C1! E 1-- =! 5 K> (2 1-- =! > 1 6+L (2 1--=! 5 K> ' 7 9& +9& +95+ +7 +9& < 57& +93 +9+; +9&7 55 +9+6 +9 ' * *? *! "#$ %! % # &7; * 70 8 ( = ' 3 C== 2D"
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
4"6"+,"0 @ &= 8 F 4$B * $ (2 H*! *!! E =D*9 4D HD >9 (2! 9 E = H9,F>$ * BE = H! YD Z>. ) D.4!%=! H4B>9H!D!D*9!! E *!!! H* HH * %!. 2!!!! E DBE H* =* J&! 5 K >.! 4&+( 4"6"++"+ $= 4$ $ H*! J$ /'. +!! (< B9 O D $ (< B. 4!! E H HH E.4!! &7+!! H %*=X >.4H&7+!!B $ 69+7 9 %I! 5+= >J ;+ 9! J&!=? &' N' % >. & 7 DBE!!. 4! B HC H B = 7 9 & J &>9 D E = & >9E = 5 9 ;J &&>9 H B! = 3 6>.) 94B!D \ H E =' % >. )$* 9 I!!! O!$ *!. 4 H $D*! >,A64 6 VI9 *! J&93!hK. 4"6"++"0 @ &= 8 F 4 H E * (2 $ D E =?? & N' % >.) *! DBE 9 * * (2 F.!I! 9 (2 H %H* *! * YD Z H* *. 4 H! H!.4/'H* =k+9k&> H! E $D* H* =? & N' % >. 4 /'! * F = +97 +93K&%H* *>.! "#$ %! % # &&
++ 1BB% B $" $ >= ( C +,, D A <A > ( C 0 DI+,, 7++ ++ 5++ ++ &++ + + +97 & &97 97 5 597 97 7 C 4 ID ' < '? 1=H.-.D!B> 1*! J(<.=6 > 1-'7 J(<.=6 >! J(<.= &;;6> 1-- = >! = > 2 07 8 $A = ( 5,A64 @ 6 4 I 4$B * (2* $D* /'* $D* E D $ E! % H* * E$H J! E *$ 9$ =' % >. 4$ H! H *! E H! * * F! &+L +97J &9&!hK=' % >.4 * J * *!! *H*. 4"6"++"4 %== % 4 *! 9! E$ $ * $F 9 \ =+97!hK>9$*! $ (2 * HBE * D F. $* * (2 F % H* *9! E$J+97! 5 K9 (2!* D!%!!."!I! =? & N' % >. 9! $! D (2 J 6+L (21--.4 DBE * 4B! $ * $! 9 $ $ * $! =1--> * =+9&;!hK>. = =A > / = = @ &= 3) = = "! "#$ %! % # &
,!! * (' H % 2 ( DC / 75 8 % > = > $ "!9 E * D H% J+97! 5 K. &@ = ( =? " ) 9 *!* J H J O $ Q!.!!$" &+( 4"6"+0"+ $= ( 9$ $ H*! J$ /'. 4 * ( D!F H J$!!.! O 4B H $'*. /' %$ D (9J (7 D!*B =J $!!! * ( D!> ( 9 J$ H D )&. 4!! H.4! E BE \ P4B H D! J;+@=! >.4H+!!B $ 79&3 9 %I! +=D\>J &; 9! ;J&7!=? &5' N' %?>. &7 3 DBE!!. 4! 5 B HC H B = & 59 39 ; &+9 & &7>9! = >9 H B D E = 79 69 &&J &5>.) 94B!D \ H D %E =' %?>. 4 H $D*!!! *+9&!hK9 *! J93!hK.'$ 9! 9J$!*!!. 9 *! E * H * 9 * $'=! &+! 5 K9-9&;;;> * J 9! J 4B.) 9 DD!E9 J * H. 9! +9&&! 5 K &7! 5 K9! E * (2 F! =!%!!>$ +9&&7! 5 K. )$ * 9 I!! J H $! 4B! * ( D! H D 9 HH * *.! "#$ %! % # &5
4"6"+0"0 @ &= 8 F > 4 HH %F!! J = P(2H ++!] &++! >. 4 (2 F H* * F J =? 3? &5 N' %?>.4!. 4 /' H* =k +9K&> H! E $D* H* =? &5) N' %?>. 4 /'! * F 9! +9 +97K&%H* *=k&97! 5 K>. ++ " $ >= ( C +,, D A <A > ( C 0 DI+,, 7++ ++ 5++ ++ &++ + + +97 & &97 97 5 597 97 7 C 4 ID ' < '? 1 1*! J(<.=6 > 1-'7 J(<.=6 >! J(<.= &;;6> 1-- = >! = > 2 0# 8 BA = >= ( 5,A++ @ 6 4 I ( 9 (2 F D H* * J &!hk! =* D >.)!I!! 9 9 (2! *! % H* *! B!E J &! 5 KH *. 4$D*! H *!! *I!! &9J9!hK.4$B * (2* $D* /'* $D* E D $ E! %H* * E$ H J! E *$ 9 =' %?>. > 4X % F * FE$J *!. %! 9(2!! * 55+! 7+!! F J * J397!hK=? 6?! "#$ %! % # &
&5 N' %?>.4 $D** HE $D* F =!%!! +93>9 H* *. 4$B * (2* $D* /'* $D* E D $ = >! * = F >E!. 9 HH $ 9 H* *9!! &97! 5 K X! *.4 $D*=/'> =+97K&>FE$J! 5 K.4 * H!E E *. 3++ " $ >= ( C +,, D A <A > ( C 0 DI+,, ++ 7++ ++ 5++ ++ &++ + + 97 7 397 &+ &97 &7 )*=! 5 K> ' < ' 1 1*! J(<.=6 > 1-'7 J(<.=6 >! J(<.= &;;6> 1-- = >! = > 2 0; 8 B B $ ( 4 >,A++ @ +6 4 I 4$ H! H *! E H! * * F! 33L +9& J &96!hK=' %?>.4! * J * *!! *H*. 4"6"+0"4 %== % > " $B DE * (2=? &5 N' %?>E * E (2D +9!hK=F > &95!hK= H >."!! +96!5K!!!. 4 *!! (2! J 6+L (2!%$D +95! 5 K= F > &! 5 K= H >!* J!!! S J DE. $! P %!D!* 9! H9 E * H! E 9 D. 9! $! D (2 J 6+L (21--.4 DBE * 4B! "#$ %! % # &7
! $ * $! 9 $ $ * $! =1--> * =+9&;!hK>.! * (' J D E. H9!!!J H* *9! +97! 5 K +96! 5 K."!* 9 P$D* H! = ++! &++! $ 9E *! &9 9! 5 K>$!! R E$! *. H 9 * J J $ * ( D! H =!* JF > I! J&! 5 K. = =DE > = =*!B!! (? = 3)? CI+D / +C *! ;,X (?=! 5 K> / 0 C1! E 1-- =! 5 K> (2 1-- =! > 1 6+L (2 1-- =! 5 K>. &3 9& +9&; +97 +965 F. 3 &9+ +93 +95+ +953. 7;; &9+ +97 +956 +953 H +6 9& +955 &9+5 &93 *. 73 &59+ +97 79;+ 7957 * F. 55 9+ +93 59+ 597 *. &93 +97 +97+ +97 76 ( = @ & B " * 9 $B DE * (2 =? &5 N' %?>E * E (2! HOH! +97!hK= > 79! 5 K= E!!! J H* *>. 4 *!! (2! J 6+L (2!%!!! =+97! 5 K J 79;! 5 K>!* J H!!. 9! $! D (2 J 6+L (21--.4 DBE * 4B! $ * $! 9 $ $ * $! * =+9&;!hK>.! * ('J D E 9!!.! "#$ %! % # &
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
! J$!* ( D!9 D!*B.( 9 H D* 9 J$!! H D - = )&>. 4!! J D*9 H9 94D HD 9$!*!!. "$ % 9J 9 * ( D! H D. ) 9!! ** DE $ ++5 = $ > *! $! H! $ =r5.7.5.5.5>.4 $", ;K+ $ E Y!B Z E H $! *C H! $ 9 E9!I!!! 9! * J 4B 9 $ E $ 9E F E H (< B9! $ *! 4B. ' * 9$ * $! D H.!5 &3( 4"6"+4"+ $= ( 9$ $ H*! J /'. 4 F! 9 O $ * $ (.4!! D!.4!!! B J. 4 H&6+!!B $ 59 9 %I! &3= > J &6 9! 5J+!=? &' N' %,>. & & DBE!!. 4! 5B HC H B E = J 69 &>9! = &9 ; &+9 &>9 H B = 59 &5J &79 &3J &>.) 9?!D JH H E E E =' %,>. 4! *! J! =! $!D > D E!. 4 H $D*!!! *+955#5!hK9 *! J+955!hK.! "#$ %! % # &6
4"6"+4"0 @ &= 8 F 4 9!! J$ D4B 9 $ Y Z * (29 E! $ H *.4 E %E JC? 9 EHE H! D E! * E E $D* H* %HH H9 E =! 9! 9 H >HE /'! $. ) 9 * I. H* HH J$!*.4 (2 9F H* * &++!]= > &6+!] = > &++! 9 E H* $!* =? ;? & N' %,>.4 H!!! %!E H D B $D*. 4 E I!!! * E $D* % F. 4 /' H* = +95K&> H! E $D* H* =? & N' %,>. = " $ >= ( A <A > 7+ ( C 0 DI+,, ++ &7+ &++ 7+ + + +97 +97 +937 & &97 &97 &937 97 97 937 5 ' '! => 1 =+K> 1-'7 =&6 > C 4 ID <? 1* 1-- => 2 09 8 $ B $ ( 5,A44 @ 4 4 I 4$B /'* $D* E D $ E!! * D F.! "#$ %! % # &;
4$ H! H *! E H! * * F! +L +95J5!hK =' %,>. 4!! * J * *9!! H H. 4"6"+4"4 %== % 4 *! \ =+95!hK>9$*! $ (2 OJ * HBE * D E! H B =? & N' %,>. '9 $* H * (29! E$J+95! 5 K9 (2!* D!%!!.! * E E!. 9! $! D (2 J 6+L (2 1--. 4 DBE *?! $ *$ J+93!hK9! * =+95! 5 K> H *!! J* B $B.4 *!!!. 4"6"+4"5 $= ) 9 *! HOH* 9 J$ /'9 * ('. A &@ 4,, IA ( =? >" - = A >>= > % &",!! * (' H % 2 ( C /= 7# 8 % > = > $ " ) 9$ % J *E E! $D $ HH *! EE E!. 4 DB D!E * H $ E $ * J*."$9J 9! $ *=",>.!! 9 E * * $ +K9 E &K+!!.! "#$ %! % # &3+
5!! H * $ K! DBE 9 HH!D I! ^ $! $D*DBE!! ** DE." * E!D! E! %! E 9! %." HH 9J$!! 9 H D HH H $. 4"6"+5"+ / 5.7.&.&.& 'D DBE 4 * D! J I!!D!D*!D!D*. 4 * J$!D!D*=/'>!*!!!*E =)->! JE *D O!% B= > =4B >. " $!! O * $'* 9 D=! D!B> * ( D!= $ (">4B. 9!D!D*! ^ O $! -' =? - 4B >!)- JE *D. *!! HH $ = B 4B > D! H$! E I $. 4 * H YDBE Z! DBE $D* %! * E! $ HH. 4!D!D* J! $ * DBE HH!.4?( -',? 4B! $ D! $ $ = J $ DBE H H! > HH B $! H *!!. H9 * YH Z! =!! DBE H DE! $! > H $*F % D D! J. ) 9?( $I. 5.7.&.&. 'D DBE 4$B DE *! = %J * S J $!D %! >9 *E!F =H 9!!9[>9 J$ % $!. 4 *!! "#$ %! % # &3&
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
4B 9 (5 D DB E ( 9 P$ HH I!9?9(5 O!!!! DB E ( (! %. 4! ** DE *!E * $ X! E!! E*E!.* *! E $ 9 B ",! F *! * * D. E $D $", 9! ^ EH9H!!!! HH H= % > $B *E E %. 9 $ HH!! 6!!!!! '?9! +! =E!! >.) 9 HH H= % >! DE. ) 9! $ *! B J $ YB! % Z! $ - \= *!! ' $ D 2 < 9 2( % 9 (!. >. B! % *! $ % HE E!B ",! '?." $ E HE H *E %.4 HE E! % * =J E!E!fE $D*! > B *E =J % > $D!!!. )! ", V++5C * DD =O 9! J >9%D $'* =O!% B>9 J'* =O 9!JH% J 4B >9 4B! * =O J DB >9 D9!!!! H D 9 %O!%. 4"6"+5"0 = = & = (BB % 5.7.&..& 'D DBE 4! ^!D!D*! *!B *E =)->! J $! * D =D $ > % D = B9H >.4 )- H $ &!hk =!>.($ E 9 D $ 9 * D * $'*! * =+97!hK>9 * D X $. 4 )- * H 9! 597 9!hK F =$ * $! )-!!!>9 *D $.! "#$ %! % # &35
( * 9 DB E B=H 5+>E D $ FE$J7+! *$ $ +!hk= * &!hk>9 +! J B!!!!.* $'* 9 9 D $! &!KD $ 9J &97!K $."!!!H DBE 9 $! J!!! * $! = *!D! * * B $ >. H 9 $ 4B $! * D %$.! "#$ %! % # &3
7 89 9/, :,9 9 & ;6 ( &, 6 ( plage exondée hors éclusée 9-9 plage exondée hors éclusée 2 4, 8 >> P =! "#$ %! % # &37
! plage exondée hors éclusée / <+, 1 +& = >=(,9<<9 > 1,<+ 2 4, \ ] 8 >> P =! "#$ %! % # &3
5.7.&.. 'D DBE 2!D $'* = H 5&>9! $! HE! I * = J +5!,?> $ K$ B. 4$ % )? E $ $ ++. HE! $ D =H F H >9 D % H = E!!!>. D E F 9! 9 * E =!!!!!>!*. 4 HE H!JH9 E J $!*!!9 D %=! E\!>.2! 9FE$J$! 9!* * E $ *!! F H.4 J HE * H!* 9 D % =!* ++>. 9!! 9!! * = H!*!* >. Cote de la ligne d'eau au barrage d'auberives (2002) cote (m NGF) 350 300 250 200 150 100 50 0 01/01/02 31/01/02 02/03/02 01/04/02 01/05/02 31/05/02 01/07/02 31/07/02 30/08/02 29/09/02 29/10/02 28/11/02 29/12/02 TRF vie sous gravier = =^ > C%2D P = C%D repro (fraie) OBR repro (fraie) vie sous gravier colonisation zones mortes 2 4+ 8 = P C 0,,0 2D 5.7.&..5 'D DBD!E 4! E D!E =D!! > $D 5+&* ++59! *! % $ % = $ >. 4!DE U *! C = H >.( 9! $ ID E $! % F= $B >9 J$!!* DD =B $!* >9 J$ B=! % >9! "#$ %! % # &33
J $ * $'* =! %!! $'* >.( 9 $ J & &7!.)!I! HOE!! 9 $B. 4 H! D =?.5J57> * %= *.6J3&> D E. Moyenne Amplitude journalière Mois mensuelle Moyenne Max. Min Mai 10,65 2,92 4,62 0,86 Juin 13,96 4,41 7,24 2,31 Juillet** 14,07 3,55 5,27 1,45 Août 11,53 1,57 4,68 0,29 Septembre 9,56 0,70 1,73 0,29 Octobre* 8,91 0,41 1,15 0,00 7; 8 F & C_ 8 + +7 = ` 8 & 7 <D Moyenne Amplitude journalière Mois mensuelle Moyenne Max. Min Mai 8,40 0,31 0,58 0,00 Juin 11,07 0,32 0,58 0,00 Juillet 12,50 0,28 0,58 0,00 Août 12,69 0,40 2,31 0,00 Septembre 9,74 0,27 2,30 0,00 Octobre* 8,68 0,45 1,15 0,00 79 8 F & @ & $= C_ 8 + +7 =D Moyenne Amplitude journalière Mois mensuelle Moyenne Max. Min Mai 8,87 1,38 6,06 0,00 Juin 14,07 5,46 6,42 3,46 Juillet 15,59 5,16 6,18 3,78 Août 15,84 4,75 6,48 2,01 Septembre 11,61 2,67 4,05 0,57 Octobre* 9,82 1,62 3,17 0,00 #, 8 F & @ & = (BB% C_ 8 + +7 =D Moyenne Amplitude journalière Mois mensuelle Moyenne Max. Min Mai** 10,90 0,96 1,44 0,57 Juin 14,47 1,35 2,06 0,87 Juillet 16,01 1,51 2,07 0,59 Août 16,22 1,57 2,91 0,89 Septembre 11,61 1,05 2,30 0,57 Octobre* 9,27 0,78 2,30 0,29 #+ 8 F & @ & & C_ 8 + +7 = ` 8 +6 4+ D! "#$ %! % # &36
! N =@> &; &3 &7 &5 && ; ( N -% - -B 3 7 5+. &! 6! &&F 7F ;F. 5F. V +V 5. &3. &. &7. 2 40 8 3 <F A? & C 8 4, +7 = 0,,4D! N =@> &; &3 &7 &5 && ; 3 7 - -% -B 5+K+K+5 &K+7K+5 6K+7K+5 &&K+K+5 7K+K+5 +;K+3K+5 5K+3K+5 +K+6K+5 +K+6K+5 +5K+;K+5 &3K+;K+5 +&K&+K+5 &7K&+K+5 2 44 8 3 <F A? & @ & $= C 8 4, +7 = 0,,4D! "#$ %! % # &3;
! =@> &; &3 &7 &5 && ; 3 7 -B - -% 5+K+K+5 &K+7K+5 6K+7K+5 &&K+K+5 7K+K+5 +;K+3K+5 5K+3K+5 +K+6K+5 +K+6K+5 +5K+;K+5 &3K+;K+5 +&K&+K+5 &7K&+K+5 2 45 8 3 <F A? & @ & = (BB% C 8 4, +7 = 0,,4D! N =@> &; &3 &7 &5 && ; 3 7 -B - -% &7K+7K+5 ;K+7K+5 &K+K+5 K+K+5 &+K+3K+5 K+3K+5 +3K+6K+5 &K+6K+5 +K+;K+5 &6K+;K+5 +K&+K+5 &K&+K+5 2 46 8 3 <F A? & @ & & C 8 +6 +7 = 0,,4D! "#$ %! % # &6+
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
5.7.&.. 'D *E ( 9 H! E B E* % DD. ( == 4 H! $D * DD B. $!! * =&!hkj$* DD >. 4 E C J$!!* DD =!* >9 J$! DD = - >9 J$ 9 J$. ' $ % J $! DD 9 9 &;;3 &;;;! HH E JHC! BE $! $= BE!J5G>H $!!! $ 9 E! =E $!! > E! $ %E H!! J $! DD. 4! E E! J$* $'*.!! C J$!! H 4B 9 D! J$$'* 9 H J$ -. ( 9 &;;6 ++&!* E! =! *! $ HH H>9 HH!. J! BE! =!* $ > B $ =* 9** 9 F>.( J $ * $'* 9 BE!J7G9 E J H.4 * J$$'* =+97!hK> E$H* $ = &7+!> H! $. $ 9 * * $'* H $". " HH 9J 9$B HH B9 E!DE! HH C! H 9 E$ * J$$'*.4! DBE HH! 9!! (" =!%!! 3! 5 K>. ($! * % D 9J = @& (9 H!> J $ * $'* = H 4B 9 @&3>9! HH!HE %! 9 E $ %E! BE 5G 7G=H.5 53>. 9! "#$ %! % # &6
!H HH H 9 *! = 7L>9!! HH H = 6+L>.( * $ %E! BE 9 E $ H * J!." HH!* $! H HH H C* 9!H EDBD!E $ 9. ) =KD> &6+++ &7+++ &+++ ;+++ +++ 5+++ + ' 4 " 4? '"? /' &+K+3K++& (&3 &3K+3K&;;6 (&=+3K&&K;3> 2 47 8 >>=> == C &ID? = & = (BB% CD ++! =QKD> &7+ &++ 7+ + ' 4 " 4? '"? /' &+K+3K++& &3K+3K&;;6 ( (&=+3K&&K;3> 2 4# 8 == CGID? = & = (BB% CD! "#$ %! % # &65
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
+" $>>= DD & 5 7 D & 5 7 '* & 5 7 4B & 5 7 4B ( D & 5 7 4B ' & 5 7 0" %= DD & 5 7 D & 5 7 '* & 5 7 4B & 5 7 4B ( D & 5 7 4B ' & 5 7 4" = > DD & 5 7 D & 5 7 '* & 5 7 4B & 5 7 4B ( D & 5 7 4B ' & 5 7 5" )? DD & 5 7 D & 5 7 '* & 5 7 4B & 5 7 4B ( D & 5 7 4B ' & 5 7 6" = DD & 5 7 D & 5 7 '* & 5 7 4B & 5 7 4B ( D & 5 7 4B ' & 5 7 7" = F DD & 5 7 D & 5 7 '* & 5 7 4B & 5 7 4B ( D & 5 7 4B ' & 5 7 #" $? a DD? -B?* D? -B?* '*? -B?* 4B? -B?* 4B ( D? -B?* 4B '? -B?* ;" 2 a DD? -B?* D? -B?* '*? -B?* 4B? -B?* 4B ( D? -B?* 4B '? -B?* 9" ) => DD? -B?* D? -B?* '*? -B?* 4B? -B?* 4B ( D? -B?* 4B '? -B?* 2 4; 8 = = = '- F? B $= F C=F D! "#$ %! % # &67
+".= @ > DD & 5 7 D & 5 7 '* & 5 7 4B & 5 7 4B ( D & 5 7 4B ' & 5 7 0" P = @ * 4". DD & 5 7 D & 5 7 '* & 5 7 4B & 5 7 4B ( D & 5 7 4B ' & 5 7 5". = DD & 5 7 D & 5 7 '* & 5 7 4B & 5 7 4B ( D & 5 7 4B ' & 5 7 6" = =? CD DD 1 & 5 7 D 1 & 5 7 '* 1 & 5 7 4B 1 & 5 7 4B ( D 1 & 5 7 4B ' 1 & 5 7 DD ( & 5 7 D ( & 5 7 '* ( & 5 7 4B ( & 5 7 4B ( D ( & 5 7 4B ' ( & 5 7 DD /N & 5 7 D /N & 5 7 '* /N & 5 7 4B /N & 5 7 4B ( D /N & 5 7 4B ' /N & 5 7 DD <N & 5 7 D <N & 5 7 '* <N & 5 7 4B <N & 5 7 4B ( D <N & 5 7 4B ' <N & 5 7 '- DD & 5 7 D & 5 7 '* & 5 7 4B & 5 7 4B ( D & 5 7 4B ' & 5 7 2 49 8 = = = '- F? B = C=F >D! "#$ %! % # &6
4 HH!DB!E! %! J.4D $D** % * *!!!. 4 B H! J DD HD!!. 4! E * DE H & E H! J $ E =! %! >.! 9 DE E = $ E(1>. 4!fE $D* H 9 * 9! %."!* E DBE % E *!fe $D* H $D!.4! $E HH D!#. 4$B HH H DE % DE J $ HH *E H.! %!E E!# 9 HE $E E *. 4$ J HE H 9 E! $ *DE =' > 9! $!*. E E *E 9 D! $ E J! E! ='* >!* E E! J *!.! %!E 9 H HH H= 1> % D 9 E D! ",9 E BH! = DE $D*> % D $.) 9 D %!E 9 E $E* E! % D E! %! J.)$! 9 E DB*E! ++5 * H! J!. ==A @ & & 8 & & &A H @ >> < C= ( % D I @ = & > == =B= & A =A == &= >= C= =D" A = @ = C= & @ = ( %A = > D! "#$ %! % # &63
4"6"+5"4 = = & = 5.7.&.5.& 'D DBE 4! ^!D!D*=D>9!D!D*9! *!!*E =)->! J $! * D =D $ > 4B 9 $ % ' =!D*>9! D!B =D>9 4B =D>. *!! 9$*! HOH* )-E 9 J$ $ 4B =E F 9 $'* >. ) O9 )- H! +9 &9!hK! = H > $ =! $! >.( $ E 9 D $ 9 $ % * J$* E! $ 9 H 9 * $ 9 E$!!.4 )- * H * 9! 7! 5 K$ =!>9 *D 4B $. (4B 9 * =! 5 K> H!! =9! 5 K>.4! =&+! +J! 5 K>." E!!! 9)?! * $ J9! 5 K *!! D =+! >. H* * $ 9 D $! 9!! J$!! H!D.4 *! * E H!. D!!**! J$ H! * O DB E. *! E H DBE * H EE.) 9D! H= * H H 9 >9!**! 9!!* \ HB. 5.7.&.5. 'D DBE ' 4B 9 E H! 9! H!. HH 9 H Y Z C E $ HH 9! *!hkue $ H J 7595!,?9 $I = * * >.4E * J! 5 K9 H H $ 9 *!. 4E *!5K9 H =& J D F!!!>9 O! % $!! *. ( $ ++ =? >9 H F9!.4 F H! H* * H $ =V!* >9 * H H J DKF."! 9 D % =!*!* %! >9 H F E.! "#$ %! % # &66
)$! 9 * H! $. * $HH HO * D E B EE = 9! F >9 E H $!*!!9 4B =J$ $ 4B >. Durée journalière de fonctionnement des groupes de l'usine de Bouvante (2002) durée (h) 24 20 16 12 8 4 0 0 30 61 91 122 152 182 213 243 274 304 334 365 = =^ > C%2D P = C%D TRF vie sous gravier repro (fraie) OBR repro (fraie) vie sous gravier colonisation zones mortes 2 5, 8 >= = C 0,,0 2D 5.7.&.5.5 'D *E ( == (4B 9 H! J$ H 4 J $!!! 9!! =5>9! H % J$!! 9 J J $ $ = > D!*B =7> F!* $ 4B.4 H!! $D!! *! = H & >." E H &;;9 H J $!! = 5 >.! "#$ %! % # &6;
) =KD> +++ &7++ &+++ 7++ + /' '"? 4? (5 ( (7 2 5+ 8 >>=> == C &ID = & = CD = > +995! =QKD> 6+ + + + + /' '"? 4? (5 ( (7 2 50 8 == CGID = & = CD = > +995! E!H! 4B 9!! J $!!. 4 E* = H &;;> J %!! $! H! = >9!H E $!! X =!! >.' E * $D 9!! $* $++5. 4 J? B! $! *D!!9 E %E!.4 D**! E E $ * E$!*! * E H! 9! H! * DE. ( 79 HH H!!I! * H. *! $! 9 E!H! "#$ %! % # &;+
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
4B 9 E!! = > E! 5!!. 4$B DB*E J $ (B! % $ J $ HE = &.+#H.++>!! * =J$ 6!E > E*E =J $ 7 B > DE D =H FJ >. )$! 9!E E! $ * DE E!# 9! *! 4B. ) 4B 9 $!DB!E E E* J$ $ 4B E$! * %!. 4 D $D** % * E E F!B 4B.4B H! 4B. 4! E * DE H & E H! J $ E =! %! >.! 9 DE E = $ E(1>. 4!fE $D* H 9 * 9! % 94B =!! >.4$D $D* H 4B $ 4B E$!!B!."!* E DBE % E *!fe $D* H $D!.4! $E HH D!#DE $. 4$B HH H DE % DE J $ HH *E H. 4 HE E E *!# 4B 9 E!E $E. 4$ J HE H 9 E! $ *DE =' > 9!$!* J$ 4B. 4 *E E! 4B 9!=E* > CE!!B J* D9! $ 4B.! %!E 9 H* HH H= 1>4B D9E D! ",9E BH! = DE $D*> % D $. 9 D %!E = (>9 $ =/$> $ $E* =<$> 4B =! % $D*>. )$! 9 E DB*E! ++5 4B H! J! 9! ++59 $ 4B = O D!! > E I * =H* c>j$* $.! "#$ %! % # &;
4"6"+5"5 = = = 5.7.&..& 'D DBE * $!! 4B 9 D. HH 9 * D %$ 9 $ 4B $! H = &J7 % $ >. 4 4B!*! J!!! J $ * $'* =! *D >. 5.7.&.. 'D *E E!! J$! * * DE I * J$!! 4B J'* U$! $!*. )$! 9 * H * $", J '* 9 E $", D %!.) 9$B!! J % HH!. 4"6"+5"6 = & = 2 C= D ( $ 9 H J$! )?=! )75&>! $ $' 4E J(. 4! H! $D!! = HH H *!? >. 4!! % 9E $H! 9!!?.( %9$D $! $ HH H9E! *!. 4 D*!!9!! %. 4$H!! HH J! * C D 9 E $ 9[ 4$!! E $D? $".! "#$ %! % # &;5
! =QKD> 7 + &7 &+ 7 + 4 /' "? ( H 2 54 8 >>=> == C &ID? = & = C2D < +994! =QKD> 7 + &7 &+ 7 + 4 /' "? ( H 2 55 8 == CGID? = & = C2D < +994? 4"6"+6"+ / 2! $ E DBE $D*!D! )7(=!DE F ++5>.!D! $ E DBE $ J$ EH EHC * 9D H9*!DE =!! "#$ %! % # &;
HH >. E %OC * DD J B9 4B * $ 4B J( D! B=O!J >. ) %!! $ E $D*DBE * < $'* 9E HD* % =HD D % >. 4"6"+6"0 % ( 9! * DD - OD! C! $ * DD! DD = &>9! DD J = >9 - = 5>. (4B 9 E OE H EH$*F $ B EDBE % EE.4! E O C? B J $ FE$ )76=4B &>9 )76FE$ )3+=4B >9 )3+FE$ $'9=7! D >J$ )3=4B 5>9 $'FE$ $ 4B =4B >. 4! O $! * DB!DE = > $. ( 4B! F B! HD* = $'9 $ 4B >. 4$ I H * YH ZY*H Z EDBE!! * E. H9 $* H B! H 9 $ I J $ = * >. DE! H 7 'JU ='> HJ H!!! J!.! "#$ %! % # &;7
) = $= * ( C*(D /&&& ';+ &5+ ( #+JG+ j=/g'>%%8 5+++ ' M7+ ' M7 ' M7 (! G&+KG+ ' M7++ +7+ 5 ;7 WE* &+KG&+ 57++#7++ 7 6+ 5 55; W 7K&+ &7++#57++ 5 ) &6 ) && 5 ) &55? +K7 ) ++#&7++ k& k&& k& (! HH *9 $D k++ #5 8 = & 4 D E! =D9 9 9 *>E EDBE * * H. L ) CI +++D = CI 9,D $= CI +4,D CB7, @ 5,D T * = P 40 & 693 7+9+ &96 +97 &97 &7 40 +9+ +9+ 395 &+97 &97 + 40 5 79+ 5796 &597 596 +967 ;3 40 39+ 7&9+ +96 &;96 &9++ &&; 2 & 579+ 9+ 569+ 69+ &97 566 2 5&9+ 7&9+ 9+ &&97 &9++ 5+6 2 5 &93 9; 69+ 393 &97 5+5& #6 8 %=> >> = * = P C LD 7+++ 7++ +++ 2 & 57++ ( EDBE 5+++ 7++ +++ 40 & 40 2 2 5 &7++ &+++ 40 7++ + 40 5 2 56 8 % = >> L! "#$ %! % # &;
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
( @ P = 8 4 J N * N'* 9 B * H. 4! * &! D. 4 D D DE * N 5+!. 2 B! D! N! *! * J! * N =! N J N >. B! N H!. 4N! =!> H J N * E E! R s! s *. 2 H =* >!! *. ( + 0 C * D % $= C '! 9!* H N! sdbe se N * J N H! E N!!. 4N =J N> * s* s E! N % J N * =D >. 9!*!! * J HD J! *!. ( 4 C ( 5C N 4 H9!! N!*!!9! N E J! HB I *!! E (!!!. ) 9 H 9 D N! N *! E!! H! = % N N!>. 4!* *!!9 E HB J N! =D! sd!s>9 N!!! E. 4 J N H N HH J N N B! J =!>!E!. " I B *E =!* 9 > J N * 9 HB B. 2 57 8 $= @ &! "#$ %! % # &;6
)$* && 9 $!* 9 J$$ $I H H!!."!* E JD! $ E J$ 9* HH $ H!.4$ I $ %! % H$.) 9 *! $ 4B 9 $'9 *!E.4$$ HD! = J> D D = 5 7!>9E! E! $ $$. ' E$ % 9 E E \ HB % 9!!! *H* J HE$J$!*!!. 5 = $ 0, 8 = 5"+ ' &> = 4, $%/BE )D$)?!!!!!!! DBE =! (HH (V /BE ((/>.! E E *$ 9 $ 9 * 9$!!! ((/U! E %. $ +++9$H!,/$)? C * 5+++ HHD =! 9 9!> 9 $!*D &! H! *9! * %E J 9 $ $ )J$ )!!! SD $ 9 &5 $H!* % 9 F J( ( 9 $ E? D D!)9! %$ $ OJ$ *!- %. && * =(,'/#,'0>! "#$ %! % # &;;
@A A< &9 (, 9 & : ( < 2 5# 8 =! "#$ %! % # ++
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